Calidris pugnax
Ruff
GC content
43.66%
Whole genome
Non-coding
1,446 bp
D-loop 552 bp
Where Ruff sits among 2,335 species
Whole-genome GC content against maximum lifespan. Ruff has the 1,318th longest lifespan of them.
- This species
- Other birds
- Other classes
Base composition
Whole genome16,902 bp
- GC skew (G − C) / (G + C)
- −0.372
- AT skew (A − T) / (A + T)
- +0.118
Life history
- Maximum lifespan
- 13.9 years
- Adult weight
- 136 g
- Female sexual maturity
- 2 years
- Male sexual maturity
- 2 years
- Postnatal growth rate
- 0.134 per day
Lifespan from AnAge · AnAge record · other traits from AnAge build 15 (HAGRID 00551)
About this record
- Lifespan source
- AnAge build 15
- Based on
- an animal in the wild
- Data quality
- acceptable, medium sample
- Licence
- Free to use, subject to the Human Ageing Genomic Resources conditions of use
- Status
- New in build 2
- In the database since
- build 2 (2026-11-13)
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome16,902 bp
Protein-coding11,376 bp
12S rRNA978 bp
16S rRNA1,570 bp
All tRNA1,545 bp
D-loop552 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 978 bp | 44.48% | −0.379 | −0.028 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,039 bp | 41.58% | −0.509 | +0.091 | |||||
| COX1Cytochrome c oxidase subunit I | 1,551 bp | 45.65% | −0.285 | −0.001 | |||||
| COX2Cytochrome c oxidase subunit II | 684 bp | 43.42% | −0.340 | +0.096 | |||||
| ATP8ATP synthase F0 subunit 8 | 168 bp | 39.88% | −0.761 | +0.188 | |||||
| ATP6ATP synthase F0 subunit 6 | 684 bp | 45.47% | −0.550 | −0.008 | |||||
| COX3Cytochrome c oxidase subunit III | 784 bp | 45.54% | −0.333 | +0.059 | |||||
| ND3NADH dehydrogenase subunit 3 | 351 bp | 45.01% | −0.468 | +0.088 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 45.12% | −0.418 | +0.055 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,378 bp | 42.96% | −0.537 | +0.130 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,815 bp | 43.69% | −0.483 | +0.159 | |||||
| ND6NADH dehydrogenase subunit 6 | 522 bp | 44.06% | −0.583 | +0.521 | |||||
| CYTBCytochrome b | 1,143 bp | 45.84% | −0.439 | +0.031 | |||||
| 12S rRNA12S ribosomal RNA | 978 bp | 46.22% | −0.155 | +0.175 | |||||
| 16S rRNA16S ribosomal RNA | 1,570 bp | 42.68% | −0.143 | +0.213 | |||||
| D-loopControl region (D-loop) | 552 bp | 44.57% | −0.138 | −0.203 |
Codon usage
Counts per codon, darker is more frequent3,797 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F5.8%
UUU60
UUC159
Leu L17.5%
UUA111
UUG16
CUU66
CUC135
CUA294
CUG41
Ile I7.7%
AUU109
AUC182
Met M4.5%
AUA136
AUG35
Val V4.5%
GUU43
GUC44
GUA64
GUG21
Ser S7.4%
UCU50
UCC75
UCA103
UCG1
AGU9
AGC44
Pro P5.7%
CCU38
CCC59
CCA113
CCG7
Thr T9.3%
ACU76
ACC146
ACA127
ACG6
Ala A7.7%
GCU70
GCC122
GCA95
GCG6
Tyr Y3.0%
UAU33
UAC82
His H2.9%
CAU25
CAC84
Gln Q2.4%
CAA84
CAG8
Asn N3.5%
AAU35
AAC98
Lys K2.3%
AAA75
AAG12
Asp D1.7%
GAU26
GAC39
Glu E2.5%
GAA82
GAG12
Cys C0.8%
UGU7
UGC24
Trp W2.8%
UGA97
UGG10
Arg R1.8%
CGU9
CGC22
CGA37
CGG1
Gly G5.8%
GGU31
GGC65
GGA95
GGG31
Stop 0.3%
UAA6
UAG1
AGA1
AGG2
Genome map
38 annotated features in NC_046877.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (43.7%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 71 | 71 bp | + | – |
| 12S rRNA | Ribosomal RNA | 71 | 1,048 | 978 bp | + | 46.2% |
| tRNA-Val | Transfer RNA | 1,048 | 1,119 | 72 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,146 | 2,715 | 1,570 bp | + | 42.7% |
| tRNA-Leu (UUR) | Transfer RNA | 2,716 | 2,789 | 74 bp | + | – |
| ND1 | Protein-coding gene | 2,806 | 3,783 | 978 bp | + | 44.5% |
| tRNA-Ile | Transfer RNA | 3,782 | 3,853 | 72 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,867 | 3,937 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 3,937 | 4,005 | 69 bp | + | – |
| ND2 | Protein-coding gene | 4,006 | 5,044 | 1,039 bp | + | 41.6% |
| tRNA-Trp | Transfer RNA | 5,045 | 5,113 | 69 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,115 | 5,183 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,186 | 5,258 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,261 | 5,327 | 67 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,327 | 5,397 | 71 bp | − | – |
| COX1 | Protein-coding gene | 5,399 | 6,949 | 1,551 bp | + | 45.7% |
| tRNA-Ser (UCN) | Transfer RNA | 6,941 | 7,014 | 74 bp | − | – |
| tRNA-Asp | Transfer RNA | 7,017 | 7,085 | 69 bp | + | – |
| COX2 | Protein-coding gene | 7,087 | 7,770 | 684 bp | + | 43.4% |
| tRNA-Lys | Transfer RNA | 7,772 | 7,842 | 71 bp | + | – |
| ATP8 | Protein-coding gene | 7,844 | 8,011 | 168 bp | + | 39.9% |
| ATP6 | Protein-coding gene | 8,002 | 8,685 | 684 bp | + | 45.5% |
| COX3 | Protein-coding gene | 8,685 | 9,468 | 784 bp | + | 45.5% |
| tRNA-Gly | Transfer RNA | 9,469 | 9,537 | 69 bp | + | – |
| ND3 | Protein-coding gene | 9,538 | 9,889 | 352 bp | + | 45.0% |
| tRNA-Arg | Transfer RNA | 9,895 | 9,963 | 69 bp | + | – |
| ND4L | Protein-coding gene | 9,965 | 10,261 | 297 bp | + | 45.1% |
| ND4 | Protein-coding gene | 10,255 | 11,632 | 1,378 bp | + | 43.0% |
| tRNA-His | Transfer RNA | 11,633 | 11,701 | 69 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,702 | 11,767 | 66 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,767 | 11,837 | 71 bp | + | – |
| ND5 | Protein-coding gene | 11,838 | 13,652 | 1,815 bp | + | 43.7% |
| CYTB | Protein-coding gene | 13,664 | 14,806 | 1,143 bp | + | 45.8% |
| tRNA-Thr | Transfer RNA | 14,809 | 14,878 | 70 bp | + | – |
| tRNA-Pro | Transfer RNA | 14,893 | 14,962 | 70 bp | − | – |
| ND6 | Protein-coding gene | 14,977 | 15,498 | 522 bp | − | 44.1% |
| tRNA-Glu | Transfer RNA | 15,502 | 15,573 | 72 bp | − | – |
| D-loop | Control region | 15,949 | 16,500 | 552 bp | – | 44.6% |