Canis lupus
Gray wolf
GC content
39.68%
Whole genome
Non-coding
1,348 bp
D-loop 1,269 bp
Where Gray wolf sits among 2,335 species
Whole-genome GC content against maximum lifespan. Gray wolf has the 870th longest lifespan of them.
- This species
- Other mammals
- Other classes
Base composition
Whole genome16,729 bp
- GC skew (G − C) / (G + C)
- −0.287
- AT skew (A − T) / (A + T)
- +0.048
Life history
- Maximum lifespan
- 20.6 years
- Adult weight
- 26.6 kg
- Birth weight
- 450 g
- Body temperature
- 38.3 °C
- Female sexual maturity
- 669 days
- Male sexual maturity
- 669 days
- Gestation or incubation
- 62 days
- Litter or clutch size
- 6
- Litters per year
- 0.8
- Postnatal growth rate
- 0.0177 per day
Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 02075)
About this record
- Lifespan source
- AnAge build 15
- Based on
- an animal in captivity
- Data quality
- high, large sample
- Licence
- Free to use, subject to the Human Ageing Genomic Resources conditions of use
- Status
- Updated in build 2
- In the database since
- build 1 (2015-07-01)
- Last changed
- build 2 (2026-11-13)
- A match to a lifespan source is under review and was not used.
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome16,729 bp
Protein-coding11,341 bp
12S rRNA954 bp
16S rRNA1,580 bp
All tRNA1,516 bp
D-loop1,269 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 957 bp | 40.96% | −0.429 | +0.012 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,044 bp | 36.69% | −0.504 | +0.123 | |||||
| COX1Cytochrome c oxidase subunit I | 1,545 bp | 40.71% | −0.126 | −0.092 | |||||
| COX2Cytochrome c oxidase subunit II | 684 bp | 39.47% | −0.304 | +0.087 | |||||
| ATP8ATP synthase F0 subunit 8 | 204 bp | 35.29% | −0.528 | +0.076 | |||||
| ATP6ATP synthase F0 subunit 6 | 681 bp | 40.24% | −0.394 | −0.027 | |||||
| COX3Cytochrome c oxidase subunit III | 784 bp | 43.11% | −0.249 | −0.090 | |||||
| ND3NADH dehydrogenase subunit 3 | 346 bp | 41.91% | −0.338 | +0.025 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 34.68% | −0.262 | −0.052 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,378 bp | 38.53% | −0.439 | +0.018 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,821 bp | 39.70% | −0.405 | +0.022 | |||||
| ND6NADH dehydrogenase subunit 6 | 528 bp | 37.88% | −0.460 | +0.287 | |||||
| CYTBCytochrome b | 1,140 bp | 41.84% | −0.338 | +0.002 | |||||
| 12S rRNA12S ribosomal RNA | 954 bp | 39.62% | −0.116 | +0.201 | |||||
| 16S rRNA16S ribosomal RNA | 1,580 bp | 38.48% | −0.095 | +0.187 | |||||
| D-loopControl region (D-loop) | 1,269 bp | 43.87% | −0.229 | −0.028 |
Codon usage
Counts per codon, darker is more frequent3,802 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.2%
UUU109
UUC126
Leu L15.6%
UUA117
UUG27
CUU95
CUC91
CUA231
CUG33
Ile I9.0%
AUU175
AUC168
Met M6.5%
AUA202
AUG45
Val V4.9%
GUU47
GUC28
GUA86
GUG24
Ser S7.4%
UCU68
UCC75
UCA76
UCG13
AGU17
AGC33
Pro P5.1%
CCU64
CCC62
CCA60
CCG7
Thr T8.0%
ACU76
ACC90
ACA120
ACG18
Ala A6.6%
GCU61
GCC87
GCA91
GCG12
Tyr Y3.8%
UAU69
UAC75
His H2.5%
CAU38
CAC57
Gln Q2.3%
CAA69
CAG20
Asn N4.1%
AAU65
AAC92
Lys K2.6%
AAA80
AAG20
Asp D1.8%
GAU29
GAC40
Glu E2.5%
GAA75
GAG19
Cys C0.7%
UGU4
UGC21
Trp W2.7%
UGA93
UGG11
Arg R1.7%
CGU6
CGC10
CGA45
CGG4
Gly G5.7%
GGU43
GGC53
GGA93
GGG27
Stop 0.3%
UAA8
UAG1
AGA1
AGG0
Genome map
38 annotated features in NC_008092.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (39.7%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 69 | 69 bp | + | – |
| 12S rRNA | Ribosomal RNA | 70 | 1,023 | 954 bp | + | 39.6% |
| tRNA-Val | Transfer RNA | 1,024 | 1,090 | 67 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,091 | 2,670 | 1,580 bp | + | 38.5% |
| tRNA-Leu (UUR) | Transfer RNA | 2,671 | 2,745 | 75 bp | + | – |
| ND1 | Protein-coding gene | 2,748 | 3,704 | 957 bp | + | 41.0% |
| tRNA-Ile | Transfer RNA | 3,704 | 3,772 | 69 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,769 | 3,843 | 75 bp | − | – |
| tRNA-Met | Transfer RNA | 3,845 | 3,914 | 70 bp | + | – |
| ND2 | Protein-coding gene | 3,915 | 4,958 | 1,044 bp | + | 36.7% |
| tRNA-Trp | Transfer RNA | 4,957 | 5,024 | 68 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,038 | 5,106 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,108 | 5,179 | 72 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,213 | 5,280 | 68 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,281 | 5,348 | 68 bp | − | – |
| COX1 | Protein-coding gene | 5,350 | 6,894 | 1,545 bp | + | 40.7% |
| tRNA-Ser (UCN) | Transfer RNA | 6,892 | 6,962 | 71 bp | − | – |
| tRNA-Asp | Transfer RNA | 6,967 | 7,034 | 68 bp | + | – |
| COX2 | Protein-coding gene | 7,035 | 7,718 | 684 bp | + | 39.5% |
| tRNA-Lys | Transfer RNA | 7,736 | 7,802 | 67 bp | + | – |
| ATP8 | Protein-coding gene | 7,804 | 8,007 | 204 bp | + | 35.3% |
| ATP6 | Protein-coding gene | 7,965 | 8,645 | 681 bp | + | 40.2% |
| COX3 | Protein-coding gene | 8,645 | 9,428 | 784 bp | + | 43.1% |
| tRNA-Gly | Transfer RNA | 9,429 | 9,496 | 68 bp | + | – |
| ND3 | Protein-coding gene | 9,497 | 9,842 | 346 bp | + | 41.9% |
| tRNA-Arg | Transfer RNA | 9,843 | 9,911 | 69 bp | + | – |
| ND4L | Protein-coding gene | 9,914 | 10,210 | 297 bp | + | 34.7% |
| ND4 | Protein-coding gene | 10,204 | 11,581 | 1,378 bp | + | 38.5% |
| tRNA-His | Transfer RNA | 11,580 | 11,650 | 71 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,651 | 11,710 | 60 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,711 | 11,780 | 70 bp | + | – |
| ND5 | Protein-coding gene | 11,781 | 13,601 | 1,821 bp | + | 39.7% |
| ND6 | Protein-coding gene | 13,585 | 14,112 | 528 bp | − | 37.9% |
| tRNA-Glu | Transfer RNA | 14,111 | 14,181 | 71 bp | − | – |
| CYTB | Protein-coding gene | 14,186 | 15,325 | 1,140 bp | + | 41.8% |
| tRNA-Thr | Transfer RNA | 15,326 | 15,395 | 70 bp | + | – |
| tRNA-Pro | Transfer RNA | 15,395 | 15,460 | 66 bp | − | – |
| D-loop | Control region | 15,461 | 16,729 | 1,269 bp | – | 43.9% |