Ciconia nigra

Black stork

Maximum lifespan

31.3 yr

AnAge record

mtDNA size

17,795 bp

RefSeq NC_023946.1

GC content

45.70%

Whole genome

Non-coding

2,298 bp

D-loop 2,206 bp

Where Black stork sits among 2,335 species

Whole-genome GC content against maximum lifespan. Black stork has the 409th longest lifespan of them.

This species
Other birds
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome17,795 bp

GC skew (G − C) / (G + C): −0.382
AT skew (A − T) / (A + T): +0.133

Life history

Maximum lifespan: 31.3 years
Adult weight: 2.9 kg
Female sexual maturity: 3 years

Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 00575)

About this record

Lifespan source: AnAge build 15
Based on: an animal in captivity
Data quality: acceptable, medium sample
Licence: Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status: Updated in build 2
In the database since: build 1 (2015-07-01)
Last changed: build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome17,795 bp

Protein-coding11,375 bp

12S rRNA971 bp

16S rRNA1,606 bp

All tRNA1,556 bp

D-loop2,206 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Ciconia nigra genes

ND1NADH dehydrogenase subunit 1	978 bp	28.5%	26.1%	12.9%	32.5%	45.40%	−0.432	+0.045
ND2NADH dehydrogenase subunit 2	1,040 bp	32.7%	23.0%	9.4%	34.9%	44.33%	−0.575	+0.174
COX1Cytochrome c oxidase subunit I	1,551 bp	27.6%	25.1%	16.1%	31.3%	47.32%	−0.322	+0.048
COX2Cytochrome c oxidase subunit II	685 bp	30.4%	24.2%	14.6%	30.8%	45.40%	−0.357	+0.112
ATP8ATP synthase F0 subunit 8	168 bp	32.7%	21.4%	5.4%	40.5%	45.83%	−0.766	+0.209
ATP6ATP synthase F0 subunit 6	684 bp	29.7%	23.4%	10.4%	36.5%	46.93%	−0.558	+0.118
COX3Cytochrome c oxidase subunit III	784 bp	27.9%	23.9%	14.9%	33.3%	48.21%	−0.381	+0.079
ND3NADH dehydrogenase subunit 3	351 bp	28.5%	23.9%	11.4%	36.2%	47.58%	−0.521	+0.087
ND4LNADH dehydrogenase subunit 4L	297 bp	26.3%	24.6%	13.8%	35.4%	49.16%	−0.438	+0.033
ND4NADH dehydrogenase subunit 4	1,378 bp	30.6%	21.7%	9.8%	37.9%	47.68%	−0.589	+0.171
ND5NADH dehydrogenase subunit 5	1,812 bp	31.2%	21.2%	11.3%	36.3%	47.57%	−0.524	+0.192
ND6NADH dehydrogenase subunit 6	522 bp	38.1%	10.9%	12.3%	38.7%	50.96%	−0.519	+0.555
CYTBCytochrome b	1,143 bp	25.0%	23.2%	13.8%	38.0%	51.79%	−0.466	+0.038
12S rRNA12S ribosomal RNA	971 bp	31.0%	19.3%	20.5%	29.2%	49.74%	−0.176	+0.234
16S rRNA16S ribosomal RNA	1,606 bp	34.1%	19.7%	18.7%	27.5%	46.20%	−0.191	+0.269
D-loopControl region (D-loop)	2,206 bp	32.4%	32.0%	15.3%	20.3%	35.59%	−0.141	+0.006

Codon usage

Counts per codon, darker is more frequent

Gene

3,796 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F5.5%

UUU52

UUC157

Leu L18.1%

UUA63

UUG24

CUU59

CUC194

CUA309

CUG39

Ile I7.6%

AUU53

AUC237

Met M4.1%

AUA130

AUG27

Val V4.3%

GUU31

GUC47

GUA68

GUG18

Ser S7.2%

UCU25

UCC107

UCA75

UCG10

AGU5

AGC52

Pro P6.0%

CCU32

CCC86

CCA105

CCG5

Thr T9.7%

ACU49

ACC161

ACA151

ACG6

Ala A7.9%

GCU39

GCC157

GCA95

GCG9

Tyr Y3.0%

UAU19

UAC96

His H2.8%

CAU20

CAC85

Gln Q2.6%

CAA90

CAG10

Asn N3.3%

AAU20

AAC105

Lys K2.2%

AAA75

AAG8

Asp D1.7%

GAU18

GAC45

Glu E2.5%

GAA81

GAG13

Cys C0.7%

UGU6

UGC22

Trp W2.8%

UGA92

UGG14

Arg R1.8%

CGU7

CGC17

CGA42

CGG4

Gly G5.8%

GGU21

GGC81

GGA85

GGG34

Stop 0.2%

UAA5

UAG1

AGA1

AGG2

Genome map

38 annotated features in NC_023946.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA22
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (45.7%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 38 features as a table

Annotated features of Ciconia nigra (NC_023946.1)
Feature	Type	Start	End	Length	Strand	GC
D-loop	Control region	1	2,206	2,206 bp	–	35.6%
tRNA-Phe	Transfer RNA	2,207	2,275	69 bp	+	–
12S rRNA	Ribosomal RNA	2,276	3,246	971 bp	+	49.7%
tRNA-Val	Transfer RNA	3,247	3,317	71 bp	+	–
16S rRNA	Ribosomal RNA	3,318	4,923	1,606 bp	+	46.2%
tRNA-Leu (UUR)	Transfer RNA	4,924	4,998	75 bp	+	–
ND1	Protein-coding gene	5,020	5,997	978 bp	+	45.4%
tRNA-Ile	Transfer RNA	5,996	6,067	72 bp	+	–
tRNA-Gln	Transfer RNA	6,076	6,146	71 bp	−	–
tRNA-Met	Transfer RNA	6,146	6,214	69 bp	+	–
ND2	Protein-coding gene	6,215	7,254	1,040 bp	+	44.3%
tRNA-Trp	Transfer RNA	7,255	7,329	75 bp	+	–
tRNA-Ala	Transfer RNA	7,331	7,399	69 bp	−	–
tRNA-Asn	Transfer RNA	7,402	7,474	73 bp	−	–
tRNA-Cys	Transfer RNA	7,477	7,543	67 bp	−	–
tRNA-Tyr	Transfer RNA	7,543	7,613	71 bp	−	–
COX1	Protein-coding gene	7,615	9,165	1,551 bp	+	47.3%
tRNA-Ser (UCN)	Transfer RNA	9,157	9,230	74 bp	−	–
tRNA-Asp	Transfer RNA	9,233	9,301	69 bp	+	–
COX2	Protein-coding gene	9,303	9,987	685 bp	+	45.4%
tRNA-Lys	Transfer RNA	9,988	10,059	72 bp	+	–
ATP8	Protein-coding gene	10,061	10,228	168 bp	+	45.8%
ATP6	Protein-coding gene	10,219	10,902	684 bp	+	46.9%
COX3	Protein-coding gene	10,902	11,685	784 bp	+	48.2%
tRNA-Gly	Transfer RNA	11,686	11,754	69 bp	+	–
ND3	Protein-coding gene	11,755	12,106	352 bp	+	47.6%
tRNA-Arg	Transfer RNA	12,111	12,181	71 bp	+	–
ND4L	Protein-coding gene	12,183	12,479	297 bp	+	49.2%
ND4	Protein-coding gene	12,473	13,850	1,378 bp	+	47.7%
tRNA-His	Transfer RNA	13,851	13,920	70 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	13,922	13,985	64 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	13,986	14,058	73 bp	+	–
ND5	Protein-coding gene	14,059	15,870	1,812 bp	+	47.6%
CYTB	Protein-coding gene	15,881	17,023	1,143 bp	+	51.8%
tRNA-Thr	Transfer RNA	17,026	17,094	69 bp	+	–
tRNA-Pro	Transfer RNA	17,107	17,176	70 bp	−	–
ND6	Protein-coding gene	17,195	17,716	522 bp	−	51.0%
tRNA-Glu	Transfer RNA	17,721	17,795	75 bp	−	–