Cottus klamathensis
Marbled sculpin
GC content
45.68%
Whole genome
Non-coding
957 bp
D-loop 521 bp
Where Marbled sculpin sits among 2,335 species
Whole-genome GC content against maximum lifespan. Marbled sculpin has the 2,036th longest lifespan of them.
- This species
- Other ray-finned fishes
- Other classes
Base composition
Whole genome16,529 bp
- GC skew (G − C) / (G + C)
- −0.282
- AT skew (A − T) / (A + T)
- +0.022
About this record
- Lifespan source
- FishBase v26.06
- Based on
- an animal in the wild
- Data quality
- acceptable
- Licence
- CC BY-NC 4.0
- Status
- New in build 2
- In the database since
- build 2 (2026-11-13)
The lifespan value is for non-commercial use only; the genome data is not restricted by it.
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome16,529 bp
Protein-coding11,406 bp
12S rRNA949 bp
16S rRNA1,664 bp
All tRNA1,553 bp
D-loop521 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 975 bp | 47.49% | −0.369 | −0.113 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,045 bp | 47.08% | −0.467 | +0.034 | |||||
| COX1Cytochrome c oxidase subunit I | 1,551 bp | 46.68% | −0.227 | −0.120 | |||||
| COX2Cytochrome c oxidase subunit II | 691 bp | 46.17% | −0.310 | +0.016 | |||||
| ATP8ATP synthase F0 subunit 8 | 168 bp | 46.43% | −0.462 | +0.133 | |||||
| ATP6ATP synthase F0 subunit 6 | 684 bp | 45.61% | −0.429 | −0.070 | |||||
| COX3Cytochrome c oxidase subunit III | 785 bp | 46.37% | −0.302 | −0.069 | |||||
| ND3NADH dehydrogenase subunit 3 | 349 bp | 46.99% | −0.341 | −0.189 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 49.50% | −0.361 | −0.147 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,381 bp | 44.53% | −0.356 | −0.052 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,839 bp | 43.83% | −0.400 | +0.005 | |||||
| ND6NADH dehydrogenase subunit 6 | 522 bp | 44.44% | −0.440 | +0.441 | |||||
| CYTBCytochrome b | 1,141 bp | 46.54% | −0.352 | −0.085 | |||||
| 12S rRNA12S ribosomal RNA | 949 bp | 47.42% | −0.102 | +0.214 | |||||
| 16S rRNA16S ribosomal RNA | 1,664 bp | 45.67% | −0.082 | +0.184 | |||||
| D-loopControl region (D-loop) | 521 bp | 44.34% | −0.100 | −0.069 |
Codon usage
Counts per codon, darker is more frequent3,807 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.0%
UUU115
UUC115
Leu L17.3%
UUA85
UUG30
CUU181
CUC157
CUA173
CUG33
Ile I7.0%
AUU148
AUC119
Met M4.1%
AUA96
AUG59
Val V5.8%
GUU71
GUC47
GUA70
GUG34
Ser S6.6%
UCU62
UCC70
UCA54
UCG8
AGU12
AGC45
Pro P5.7%
CCU74
CCC88
CCA49
CCG5
Thr T8.4%
ACU53
ACC143
ACA115
ACG7
Ala A8.7%
GCU75
GCC171
GCA81
GCG5
Tyr Y2.8%
UAU33
UAC74
His H2.9%
CAU38
CAC74
Gln Q2.6%
CAA85
CAG15
Asn N3.1%
AAU32
AAC86
Lys K1.9%
AAA67
AAG5
Asp D1.9%
GAU20
GAC54
Glu E2.6%
GAA84
GAG15
Cys C0.7%
UGU8
UGC17
Trp W3.1%
UGA105
UGG14
Arg R2.0%
CGU15
CGC16
CGA36
CGG9
Gly G6.5%
GGU41
GGC79
GGA84
GGG44
Stop 0.2%
UAA6
UAG1
AGA0
AGG0
Genome map
38 annotated features in NC_066916.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (45.7%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 68 | 68 bp | + | – |
| 12S rRNA | Ribosomal RNA | 69 | 1,017 | 949 bp | + | 47.4% |
| tRNA-Val | Transfer RNA | 1,018 | 1,089 | 72 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,116 | 2,779 | 1,664 bp | + | 45.7% |
| tRNA-Leu (UUR) | Transfer RNA | 2,780 | 2,853 | 74 bp | + | – |
| ND1 | Protein-coding gene | 2,854 | 3,828 | 975 bp | + | 47.5% |
| tRNA-Ile | Transfer RNA | 3,833 | 3,902 | 70 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,902 | 3,972 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 3,972 | 4,041 | 70 bp | + | – |
| ND2 | Protein-coding gene | 4,042 | 5,086 | 1,045 bp | + | 47.1% |
| tRNA-Trp | Transfer RNA | 5,087 | 5,157 | 71 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,159 | 5,227 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,229 | 5,301 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,339 | 5,404 | 66 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,405 | 5,474 | 70 bp | − | – |
| COX1 | Protein-coding gene | 5,476 | 7,026 | 1,551 bp | + | 46.7% |
| tRNA-Ser (UCN) | Transfer RNA | 7,027 | 7,097 | 71 bp | − | – |
| tRNA-Asp | Transfer RNA | 7,101 | 7,173 | 73 bp | + | – |
| COX2 | Protein-coding gene | 7,191 | 7,881 | 691 bp | + | 46.2% |
| tRNA-Lys | Transfer RNA | 7,882 | 7,955 | 74 bp | + | – |
| ATP8 | Protein-coding gene | 7,957 | 8,124 | 168 bp | + | 46.4% |
| ATP6 | Protein-coding gene | 8,115 | 8,798 | 684 bp | + | 45.6% |
| COX3 | Protein-coding gene | 8,798 | 9,582 | 785 bp | + | 46.4% |
| tRNA-Gly | Transfer RNA | 9,583 | 9,655 | 73 bp | + | – |
| ND3 | Protein-coding gene | 9,656 | 10,004 | 349 bp | + | 47.0% |
| tRNA-Arg | Transfer RNA | 10,005 | 10,073 | 69 bp | + | – |
| ND4L | Protein-coding gene | 10,074 | 10,370 | 297 bp | + | 49.5% |
| ND4 | Protein-coding gene | 10,364 | 11,744 | 1,381 bp | + | 44.5% |
| tRNA-His | Transfer RNA | 11,745 | 11,813 | 69 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,814 | 11,881 | 68 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,886 | 11,958 | 73 bp | + | – |
| ND5 | Protein-coding gene | 11,959 | 13,797 | 1,839 bp | + | 43.8% |
| ND6 | Protein-coding gene | 13,794 | 14,315 | 522 bp | − | 44.4% |
| tRNA-Glu | Transfer RNA | 14,316 | 14,384 | 69 bp | − | – |
| CYTB | Protein-coding gene | 14,389 | 15,529 | 1,141 bp | + | 46.5% |
| tRNA-Thr | Transfer RNA | 15,530 | 15,601 | 72 bp | + | – |
| tRNA-Pro | Transfer RNA | 15,601 | 15,671 | 71 bp | − | – |
| D-loop | Control region | 15,957 | 16,477 | 521 bp | – | 44.3% |