Epinephelus fuscoguttatus
Brown-marbled grouper
GC content
43.90%
Whole genome
Non-coding
1,021 bp
D-loop 948 bp
Where Brown-marbled grouper sits among 2,335 species
Whole-genome GC content against maximum lifespan. Brown-marbled grouper has the 267th longest lifespan of them.
- This species
- Other ray-finned fishes
- Other classes
Base composition
Whole genome16,648 bp
- GC skew (G − C) / (G + C)
- −0.289
- AT skew (A − T) / (A + T)
- +0.040
About this record
- Lifespan source
- FishBase v26.06
- Based on
- an animal in the wild
- Data quality
- acceptable
- Licence
- CC BY-NC 4.0
- Status
- New in build 2
- In the database since
- build 2 (2026-11-13)
The lifespan value is for non-commercial use only; the genome data is not restricted by it.
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome16,648 bp
Protein-coding11,406 bp
12S rRNA952 bp
16S rRNA1,705 bp
All tRNA1,564 bp
D-loop948 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 975 bp | 46.15% | −0.418 | −0.002 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,045 bp | 45.74% | −0.506 | +0.101 | |||||
| COX1Cytochrome c oxidase subunit I | 1,551 bp | 42.94% | −0.195 | −0.121 | |||||
| COX2Cytochrome c oxidase subunit II | 691 bp | 42.40% | −0.242 | +0.030 | |||||
| ATP8ATP synthase F0 subunit 8 | 168 bp | 44.05% | −0.432 | 0.000 | |||||
| ATP6ATP synthase F0 subunit 6 | 684 bp | 42.69% | −0.411 | −0.026 | |||||
| COX3Cytochrome c oxidase subunit III | 785 bp | 45.61% | −0.291 | −0.035 | |||||
| ND3NADH dehydrogenase subunit 3 | 349 bp | 44.99% | −0.389 | −0.177 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 51.52% | −0.412 | −0.056 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,381 bp | 43.16% | −0.383 | −0.024 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,839 bp | 43.39% | −0.393 | +0.001 | |||||
| ND6NADH dehydrogenase subunit 6 | 522 bp | 46.17% | −0.378 | +0.409 | |||||
| CYTBCytochrome b | 1,141 bp | 45.93% | −0.397 | −0.028 | |||||
| 12S rRNA12S ribosomal RNA | 952 bp | 47.69% | −0.106 | +0.153 | |||||
| 16S rRNA16S ribosomal RNA | 1,705 bp | 46.22% | −0.117 | +0.243 | |||||
| D-loopControl region (D-loop) | 948 bp | 30.80% | −0.110 | +0.009 |
Codon usage
Counts per codon, darker is more frequent3,807 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.2%
UUU107
UUC130
Leu L17.3%
UUA99
UUG23
CUU159
CUC132
CUA214
CUG33
Ile I7.3%
AUU169
AUC109
Met M4.0%
AUA107
AUG44
Val V5.6%
GUU68
GUC46
GUA82
GUG18
Ser S6.4%
UCU36
UCC79
UCA70
UCG4
AGU10
AGC43
Pro P5.8%
CCU67
CCC91
CCA57
CCG5
Thr T8.1%
ACU51
ACC117
ACA134
ACG6
Ala A9.0%
GCU70
GCC139
GCA116
GCG16
Tyr Y2.9%
UAU40
UAC71
His H2.8%
CAU32
CAC75
Gln Q2.4%
CAA86
CAG7
Asn N3.1%
AAU31
AAC87
Lys K2.0%
AAA73
AAG4
Asp D2.0%
GAU22
GAC55
Glu E2.5%
GAA85
GAG12
Cys C0.8%
UGU18
UGC13
Trp W3.1%
UGA107
UGG12
Arg R2.1%
CGU11
CGC16
CGA50
CGG3
Gly G6.3%
GGU49
GGC76
GGA87
GGG27
Stop 0.2%
UAA6
UAG1
AGA0
AGG0
Genome map
38 annotated features in NC_020046.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (43.9%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 70 | 70 bp | + | – |
| 12S rRNA | Ribosomal RNA | 71 | 1,022 | 952 bp | + | 47.7% |
| tRNA-Val | Transfer RNA | 1,023 | 1,092 | 70 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,094 | 2,798 | 1,705 bp | + | 46.2% |
| tRNA-Leu (UUR) | Transfer RNA | 2,799 | 2,874 | 76 bp | + | – |
| ND1 | Protein-coding gene | 2,875 | 3,849 | 975 bp | + | 46.2% |
| tRNA-Ile | Transfer RNA | 3,854 | 3,923 | 70 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,923 | 3,993 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 3,994 | 4,062 | 69 bp | + | – |
| ND2 | Protein-coding gene | 4,063 | 5,107 | 1,045 bp | + | 45.7% |
| tRNA-Trp | Transfer RNA | 5,108 | 5,178 | 71 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,180 | 5,248 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,249 | 5,321 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,361 | 5,427 | 67 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,428 | 5,498 | 71 bp | − | – |
| COX1 | Protein-coding gene | 5,500 | 7,050 | 1,551 bp | + | 42.9% |
| tRNA-Ser (UCN) | Transfer RNA | 7,052 | 7,122 | 71 bp | − | – |
| tRNA-Asp | Transfer RNA | 7,126 | 7,198 | 73 bp | + | – |
| COX2 | Protein-coding gene | 7,207 | 7,897 | 691 bp | + | 42.4% |
| tRNA-Lys | Transfer RNA | 7,898 | 7,971 | 74 bp | + | – |
| ATP8 | Protein-coding gene | 7,973 | 8,140 | 168 bp | + | 44.1% |
| ATP6 | Protein-coding gene | 8,131 | 8,814 | 684 bp | + | 42.7% |
| COX3 | Protein-coding gene | 8,814 | 9,598 | 785 bp | + | 45.6% |
| tRNA-Gly | Transfer RNA | 9,599 | 9,670 | 72 bp | + | – |
| ND3 | Protein-coding gene | 9,671 | 10,019 | 349 bp | + | 45.0% |
| tRNA-Arg | Transfer RNA | 10,020 | 10,088 | 69 bp | + | – |
| ND4L | Protein-coding gene | 10,089 | 10,385 | 297 bp | + | 51.5% |
| ND4 | Protein-coding gene | 10,379 | 11,759 | 1,381 bp | + | 43.2% |
| tRNA-His | Transfer RNA | 11,760 | 11,829 | 70 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,830 | 11,901 | 72 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,909 | 11,981 | 73 bp | + | – |
| ND5 | Protein-coding gene | 11,982 | 13,820 | 1,839 bp | + | 43.4% |
| ND6 | Protein-coding gene | 13,817 | 14,338 | 522 bp | − | 46.2% |
| tRNA-Glu | Transfer RNA | 14,339 | 14,408 | 70 bp | − | – |
| CYTB | Protein-coding gene | 14,416 | 15,556 | 1,141 bp | + | 45.9% |
| tRNA-Thr | Transfer RNA | 15,557 | 15,630 | 74 bp | + | – |
| tRNA-Pro | Transfer RNA | 15,631 | 15,700 | 70 bp | + | – |
| D-loop | Control region | 15,701 | 16,648 | 948 bp | – | 30.8% |