Malacochersus tornieri

Pancake tortoise

Maximum lifespan

25.9 yr

AnAge record

mtDNA size

19,202 bp

RefSeq NC_007700.1

GC content

37.83%

Whole genome

Non-coding

3,678 bp

D-loop 1,683 bp

Where Pancake tortoise sits among 2,335 species

Whole-genome GC content against maximum lifespan. Pancake tortoise has the 607th longest lifespan of them.

This species
Other reptiles
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome19,202 bp

GC skew (G − C) / (G + C): −0.410
AT skew (A − T) / (A + T): +0.128

Life history

Maximum lifespan: 25.9 years
Adult weight: 391.3 g

Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 03779)

About this record

Lifespan source: AnAge build 15
Based on: an animal in captivity
Data quality: acceptable, large sample
Licence: Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status: Updated in build 2
In the database since: build 1 (2015-07-01)
Last changed: build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome19,202 bp

Protein-coding11,353 bp

12S rRNA969 bp

16S rRNA1,609 bp

All tRNA1,606 bp

D-loop1,683 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Malacochersus tornieri genes

ND1NADH dehydrogenase subunit 1	971 bp	32.7%	28.3%	10.7%	28.2%	38.93%	−0.450	+0.073
ND2NADH dehydrogenase subunit 2	1,039 bp	36.1%	25.6%	9.3%	29.0%	38.31%	−0.513	+0.170
COX1Cytochrome c oxidase subunit I	1,545 bp	30.4%	28.3%	15.3%	26.1%	41.36%	−0.261	+0.035
COX2Cytochrome c oxidase subunit II	687 bp	36.1%	25.6%	12.2%	26.1%	38.28%	−0.361	+0.170
ATP8ATP synthase F0 subunit 8	168 bp	41.7%	22.0%	6.5%	29.8%	36.31%	−0.639	+0.308
ATP6ATP synthase F0 subunit 6	684 bp	35.4%	27.2%	8.3%	29.1%	37.43%	−0.555	+0.131
COX3Cytochrome c oxidase subunit III	784 bp	31.8%	26.0%	14.2%	28.1%	42.22%	−0.329	+0.099
ND3NADH dehydrogenase subunit 3	351 bp	29.6%	30.5%	10.3%	29.6%	39.89%	−0.486	−0.014
ND4LNADH dehydrogenase subunit 4L	297 bp	30.3%	30.6%	10.4%	28.6%	39.06%	−0.466	−0.006
ND4NADH dehydrogenase subunit 4	1,378 bp	36.2%	26.4%	10.1%	27.3%	37.37%	−0.460	+0.156
ND5NADH dehydrogenase subunit 5	1,803 bp	35.8%	25.5%	9.1%	29.7%	38.77%	−0.531	+0.168
ND6NADH dehydrogenase subunit 6	525 bp	49.1%	13.7%	7.4%	29.7%	37.14%	−0.600	+0.564
CYTBCytochrome b	1,144 bp	32.8%	25.8%	10.4%	31.0%	41.43%	−0.498	+0.119
12S rRNA12S ribosomal RNA	969 bp	36.9%	21.4%	16.7%	25.0%	41.69%	−0.198	+0.267
16S rRNA16S ribosomal RNA	1,609 bp	39.5%	21.8%	16.2%	22.5%	38.72%	−0.162	+0.288
D-loopControl region (D-loop)	1,683 bp	35.2%	34.4%	8.5%	21.9%	30.42%	−0.441	+0.011

Codon usage

Counts per codon, darker is more frequent

Gene

3,790 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F5.8%

UUU88

UUC132

Leu L16.3%

UUA132

UUG19

CUU65

CUC68

CUA316

CUG19

Ile I8.3%

AUU138

AUC176

Met M6.1%

AUA204

AUG27

Val V4.0%

GUU35

GUC16

GUA88

GUG13

Ser S7.7%

UCU47

UCC68

UCA122

UCG4

AGU13

AGC36

Pro P5.3%

CCU19

CCC27

CCA152

CCG2

Thr T10.0%

ACU66

ACC144

ACA165

ACG5

Ala A6.0%

GCU39

GCC99

GCA85

GCG4

Tyr Y3.4%

UAU51

UAC77

His H2.8%

CAU45

CAC63

Gln Q2.6%

CAA97

CAG2

Asn N4.1%

AAU46

AAC111

Lys K2.4%

AAA91

AAG1

Asp D1.6%

GAU30

GAC32

Glu E2.3%

GAA81

GAG7

Cys C0.7%

UGU13

UGC14

Trp W2.9%

UGA99

UGG10

Arg R1.7%

CGU7

CGC11

CGA47

CGG1

Gly G5.6%

GGU30

GGC57

GGA106

GGG20

Stop 0.2%

UAA5

UAG1

AGA0

AGG2

Genome map

39 annotated features in NC_007700.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA23
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (37.8%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 39 features as a table

Annotated features of Malacochersus tornieri (NC_007700.1)
Feature	Type	Start	End	Length	Strand	GC
tRNA-Phe	Transfer RNA	1	69	69 bp	+	–
tRNA-Phe	Transfer RNA	2,020	2,088	69 bp	+	–
12S rRNA	Ribosomal RNA	2,087	3,055	969 bp	+	41.7%
tRNA-Val	Transfer RNA	3,056	3,127	72 bp	+	–
16S rRNA	Ribosomal RNA	3,128	4,736	1,609 bp	+	38.7%
tRNA-Leu (UUR)	Transfer RNA	4,735	4,810	76 bp	+	–
ND1	Protein-coding gene	4,811	5,781	971 bp	+	38.9%
tRNA-Ile	Transfer RNA	5,782	5,850	69 bp	+	–
tRNA-Gln	Transfer RNA	5,849	5,919	71 bp	−	–
tRNA-Met	Transfer RNA	5,919	5,988	70 bp	+	–
ND2	Protein-coding gene	5,989	7,027	1,039 bp	+	38.3%
tRNA-Trp	Transfer RNA	7,028	7,103	76 bp	+	–
tRNA-Ala	Transfer RNA	7,105	7,173	69 bp	−	–
tRNA-Asn	Transfer RNA	7,175	7,247	73 bp	−	–
tRNA-Cys	Transfer RNA	7,275	7,340	66 bp	−	–
tRNA-Tyr	Transfer RNA	7,341	7,411	71 bp	−	–
COX1	Protein-coding gene	7,413	8,957	1,545 bp	+	41.4%
tRNA-Ser (UCN)	Transfer RNA	8,949	9,019	71 bp	−	–
tRNA-Asp	Transfer RNA	9,020	9,088	69 bp	+	–
COX2	Protein-coding gene	9,089	9,775	687 bp	+	38.3%
tRNA-Lys	Transfer RNA	9,781	9,852	72 bp	+	–
ATP8	Protein-coding gene	9,854	10,021	168 bp	+	36.3%
ATP6	Protein-coding gene	10,012	10,695	684 bp	+	37.4%
COX3	Protein-coding gene	10,695	11,478	784 bp	+	42.2%
tRNA-Gly	Transfer RNA	11,479	11,545	67 bp	+	–
ND3	Protein-coding gene	11,546	11,897	352 bp	+	39.9%
tRNA-Arg	Transfer RNA	11,898	11,966	69 bp	+	–
ND4L	Protein-coding gene	11,967	12,263	297 bp	+	39.1%
ND4	Protein-coding gene	12,257	13,635	1,379 bp	+	37.4%
tRNA-His	Transfer RNA	13,636	13,703	68 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	13,704	13,767	64 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	13,767	13,838	72 bp	+	–
ND5	Protein-coding gene	13,841	15,643	1,803 bp	+	38.8%
ND6	Protein-coding gene	15,639	16,163	525 bp	−	37.1%
tRNA-Glu	Transfer RNA	16,164	16,231	68 bp	−	–
CYTB	Protein-coding gene	16,236	17,379	1,144 bp	+	41.4%
tRNA-Thr	Transfer RNA	17,380	17,450	71 bp	+	–
tRNA-Pro	Transfer RNA	17,452	17,519	68 bp	−	–
D-loop	Control region	17,520	19,202	1,683 bp	–	30.4%