MitoAge

Manouria emys

Asian brown tortoise, or East Indian upland tortoise

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Maximum lifespan
19.9 yr
AnAge record
mtDNA size
16,455 bp
RefSeq NC_007693.1
GC content
40.03%
Whole genome
Non-coding
995 bp
D-loop 948 bp

Where Asian brown tortoise, or East Indian upland tortoise sits among 2,335 species

Whole-genome GC content against maximum lifespan. Asian brown tortoise, or East Indian upland tortoise has the 953rd longest lifespan of them.

10203040501 yr10 yr100 yrMaximum lifespan, log scaleGC %Asian brown tortoise, or East Indian upland tortoise
  • This species
  • Other reptiles
  • Other classes
Plot other mtDNA features with this species highlighted

Base composition

Whole genome16,455 bp
GC skew (G − C) / (G + C)
−0.341
AT skew (A − T) / (A + T)
+0.137

Life history

Maximum lifespan
19.9 years
Adult weight
26 kg
Gestation or incubation
74 days

Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 03780)

About this record

Lifespan source
AnAge build 15
Based on
an animal in captivity
Data quality
acceptable, medium sample
Licence
Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status
Updated in build 2
In the database since
build 1 (2015-07-01)
Last changed
build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq
Whole genome16,455 bp
Protein-coding11,344 bp
12S rRNA970 bp
16S rRNA1,610 bp
All tRNA1,551 bp
D-loop948 bp

Genes and regions

Select a column to sort; select a gene to compare it across species
Composition
ND1NADH dehydrogenase subunit 1972 bp30.7%28.0%12.7%28.7%41.36%−0.388+0.046
ND2NADH dehydrogenase subunit 21,041 bp36.9%23.3%8.8%30.9%39.77%−0.556+0.225
COX1Cytochrome c oxidase subunit I1,548 bp28.4%29.7%17.2%24.7%41.93%−0.177−0.023
COX2Cytochrome c oxidase subunit II687 bp35.2%26.1%13.7%25.0%38.72%−0.293+0.150
ATP8ATP synthase F0 subunit 8168 bp39.3%25.6%6.0%29.2%35.12%−0.661+0.211
ATP6ATP synthase F0 subunit 6681 bp32.5%26.9%10.9%29.8%40.68%−0.466+0.094
COX3Cytochrome c oxidase subunit III784 bp29.2%26.7%16.2%27.9%44.13%−0.266+0.046
ND3NADH dehydrogenase subunit 3351 bp30.8%27.6%12.8%28.8%41.60%−0.384+0.054
ND4LNADH dehydrogenase subunit 4L297 bp30.0%29.3%11.4%29.3%40.74%−0.438+0.011
ND4NADH dehydrogenase subunit 41,378 bp34.7%25.3%11.3%28.7%39.99%−0.434+0.156
ND5NADH dehydrogenase subunit 51,791 bp34.7%25.7%10.5%29.0%39.53%−0.469+0.149
ND6NADH dehydrogenase subunit 6525 bp45.9%14.5%8.6%31.0%39.62%−0.567+0.521
CYTBCytochrome b1,144 bp30.7%27.0%12.0%30.3%42.31%−0.434+0.064
12S rRNA12S ribosomal RNA970 bp37.0%20.9%16.6%25.5%42.06%−0.211+0.278
16S rRNA16S ribosomal RNA1,610 bp38.3%21.5%17.1%23.2%40.25%−0.151+0.281
D-loopControl region (D-loop)948 bp34.1%34.3%11.4%20.3%31.65%−0.280−0.003

Codon usage

Counts per codon, darker is more frequent

3,788 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F5.8%
UUU87
UUC133
Leu L15.8%
UUA125
UUG18
CUU60
CUC69
CUA283
CUG42
Ile I8.3%
AUU135
AUC181
Met M6.3%
AUA194
AUG44
Val V4.2%
GUU40
GUC20
GUA86
GUG14
Ser S7.5%
UCU47
UCC63
UCA120
UCG6
AGU12
AGC36
Pro P5.5%
CCU20
CCC32
CCA143
CCG12
Thr T9.1%
ACU70
ACC125
ACA143
ACG8
Ala A6.9%
GCU58
GCC99
GCA88
GCG15
Tyr Y3.2%
UAU44
UAC77
His H2.8%
CAU41
CAC64
Gln Q2.7%
CAA96
CAG5
Asn N3.8%
AAU41
AAC104
Lys K2.4%
AAA84
AAG7
Asp D1.7%
GAU26
GAC39
Glu E2.4%
GAA86
GAG6
Cys C0.8%
UGU14
UGC15
Trp W3.0%
UGA99
UGG13
Arg R1.9%
CGU8
CGC14
CGA45
CGG5
Gly G5.7%
GGU27
GGC62
GGA92
GGG36
Stop 0.3%
UAA5
UAG3
AGA0
AGG2

Genome map

38 annotated features in NC_007693.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

  • Protein-coding13
  • rRNA2
  • tRNA22
  • D-loop1
  • Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
  • Middle ring: GC% of each gene against the whole genome (40.0%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
FeatureTypeStartEndLengthStrandGC
tRNA-PheTransfer RNA17070 bp+–
12S rRNARibosomal RNA711,040970 bp+42.1%
tRNA-ValTransfer RNA1,0411,11070 bp+–
16S rRNARibosomal RNA1,1112,7201,610 bp+40.3%
tRNA-Leu (UUR)Transfer RNA2,7212,79777 bp+–
ND1Protein-coding gene2,7993,770972 bp+41.4%
tRNA-IleTransfer RNA3,7703,83970 bp+–
tRNA-GlnTransfer RNA3,8393,90971 bp−–
tRNA-MetTransfer RNA3,9093,97769 bp+–
ND2Protein-coding gene3,9785,0181,041 bp+39.8%
tRNA-TrpTransfer RNA5,0175,09074 bp+–
tRNA-AlaTransfer RNA5,0925,16069 bp−–
tRNA-AsnTransfer RNA5,1625,23473 bp−–
tRNA-CysTransfer RNA5,2625,32665 bp−–
tRNA-TyrTransfer RNA5,3275,39872 bp−–
COX1Protein-coding gene5,4006,9471,548 bp+41.9%
tRNA-Ser (UCN)Transfer RNA6,9397,00971 bp+–
tRNA-AspTransfer RNA7,0107,07970 bp+–
COX2Protein-coding gene7,0807,766687 bp+38.7%
tRNA-LysTransfer RNA7,7717,84474 bp+–
ATP8Protein-coding gene7,8458,012168 bp+35.1%
ATP6Protein-coding gene8,0038,683681 bp+40.7%
COX3Protein-coding gene8,6839,466784 bp+44.1%
tRNA-GlyTransfer RNA9,4679,53468 bp+–
ND3Protein-coding gene9,5359,886352 bp+41.6%
tRNA-ArgTransfer RNA9,8859,95874 bp+–
ND4LProtein-coding gene9,95810,254297 bp+40.7%
ND4Protein-coding gene10,24811,6251,378 bp+40.0%
tRNA-HisTransfer RNA11,62611,69570 bp+–
tRNA-Ser (AGY)Transfer RNA11,69611,76267 bp+–
tRNA-Leu (CUN)Transfer RNA11,76211,83372 bp+–
ND5Protein-coding gene11,83413,6241,791 bp+39.5%
ND6Protein-coding gene13,62014,144525 bp−39.6%
tRNA-GluTransfer RNA14,14514,21268 bp−–
CYTBProtein-coding gene14,21715,3601,144 bp+42.3%
tRNA-ThrTransfer RNA15,36115,43171 bp+–
tRNA-ProTransfer RNA15,43915,50769 bp+–
D-loopControl region15,50816,455948 bp–31.7%

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