Parahyaena brunnea
Brown hyena
GC content
42.57%
Whole genome
Non-coding
1,820 bp
D-loop 1,758 bp
Where Brown hyena sits among 2,335 species
Whole-genome GC content against maximum lifespan. Brown hyena has the 399th longest lifespan of them.
- This species
- Other mammals
- Other classes
Base composition
Whole genome17,197 bp
- GC skew (G − C) / (G + C)
- −0.314
- AT skew (A − T) / (A + T)
- +0.129
Life history
- Maximum lifespan
- 31.9 years
- Adult weight
- 42.3 kg
- Birth weight
- 688 g
- Female sexual maturity
- 2.7 years
- Gestation or incubation
- 92 days
- Litter or clutch size
- 2.5
- Litters per year
- 0.4
Lifespan from AnAge · AnAge record · other traits from AnAge build 15 (HAGRID 02159)
About this record
- Lifespan source
- AnAge build 15
- Based on
- an animal in captivity
- Data quality
- acceptable, small sample
- Licence
- Free to use, subject to the Human Ageing Genomic Resources conditions of use
- Status
- New in build 2
- In the database since
- build 2 (2026-11-13)
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome17,197 bp
Protein-coding11,339 bp
12S rRNA959 bp
16S rRNA1,570 bp
All tRNA1,511 bp
D-loop1,758 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 956 bp | 44.77% | −0.439 | +0.125 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,042 bp | 40.79% | −0.558 | +0.254 | |||||
| COX1Cytochrome c oxidase subunit I | 1,545 bp | 45.44% | −0.168 | −0.068 | |||||
| COX2Cytochrome c oxidase subunit II | 684 bp | 43.42% | −0.279 | +0.096 | |||||
| ATP8ATP synthase F0 subunit 8 | 204 bp | 33.82% | −0.594 | +0.244 | |||||
| ATP6ATP synthase F0 subunit 6 | 681 bp | 44.20% | −0.422 | +0.058 | |||||
| COX3Cytochrome c oxidase subunit III | 784 bp | 46.81% | −0.275 | −0.007 | |||||
| ND3NADH dehydrogenase subunit 3 | 347 bp | 43.80% | −0.329 | +0.077 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 39.73% | −0.339 | −0.073 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,378 bp | 41.95% | −0.419 | +0.100 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,821 bp | 42.01% | −0.441 | +0.125 | |||||
| ND6NADH dehydrogenase subunit 6 | 528 bp | 39.77% | −0.457 | +0.327 | |||||
| CYTBCytochrome b | 1,140 bp | 44.04% | −0.375 | +0.088 | |||||
| 12S rRNA12S ribosomal RNA | 959 bp | 43.90% | −0.116 | +0.234 | |||||
| 16S rRNA16S ribosomal RNA | 1,570 bp | 40.76% | −0.119 | +0.239 | |||||
| D-loopControl region (D-loop) | 1,758 bp | 42.83% | −0.309 | +0.180 |
Codon usage
Counts per codon, darker is more frequent3,800 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.0%
UUU88
UUC139
Leu L15.8%
UUA58
UUG26
CUU51
CUC101
CUA291
CUG74
Ile I8.6%
AUU124
AUC204
Met M6.1%
AUA176
AUG56
Val V4.8%
GUU26
GUC35
GUA92
GUG30
Ser S7.3%
UCU44
UCC65
UCA107
UCG11
AGU13
AGC39
Pro P5.3%
CCU41
CCC78
CCA71
CCG12
Thr T8.4%
ACU42
ACC109
ACA157
ACG11
Ala A7.0%
GCU38
GCC128
GCA95
GCG6
Tyr Y3.6%
UAU55
UAC81
His H2.4%
CAU15
CAC78
Gln Q2.4%
CAA80
CAG13
Asn N4.1%
AAU42
AAC114
Lys K2.7%
AAA88
AAG16
Asp D1.7%
GAU19
GAC45
Glu E2.7%
GAA79
GAG22
Cys C0.6%
UGU7
UGC16
Trp W2.8%
UGA83
UGG22
Arg R1.7%
CGU4
CGC16
CGA37
CGG8
Gly G5.6%
GGU42
GGC52
GGA87
GGG33
Stop 0.2%
UAA7
UAG0
AGA1
AGG0
Genome map
38 annotated features in NC_038159.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (42.6%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 879 | 948 | 70 bp | + | – |
| 12S rRNA | Ribosomal RNA | 949 | 1,907 | 959 bp | + | 43.9% |
| tRNA-Val | Transfer RNA | 1,908 | 1,975 | 68 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,976 | 3,545 | 1,570 bp | + | 40.8% |
| tRNA-Leu (UUR) | Transfer RNA | 3,546 | 3,620 | 75 bp | + | – |
| ND1 | Protein-coding gene | 3,623 | 4,578 | 956 bp | + | 44.8% |
| tRNA-Ile | Transfer RNA | 4,579 | 4,647 | 69 bp | + | – |
| tRNA-Gln | Transfer RNA | 4,645 | 4,717 | 73 bp | − | – |
| tRNA-Met | Transfer RNA | 4,719 | 4,787 | 69 bp | + | – |
| ND2 | Protein-coding gene | 4,788 | 5,829 | 1,042 bp | + | 40.8% |
| tRNA-Trp | Transfer RNA | 5,830 | 5,897 | 68 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,912 | 5,980 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,982 | 6,054 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 6,087 | 6,154 | 68 bp | − | – |
| tRNA-Tyr | Transfer RNA | 6,155 | 6,221 | 67 bp | − | – |
| COX1 | Protein-coding gene | 6,223 | 7,767 | 1,545 bp | + | 45.4% |
| tRNA-Ser (UCN) | Transfer RNA | 7,766 | 7,835 | 70 bp | − | – |
| tRNA-Asp | Transfer RNA | 7,840 | 7,908 | 69 bp | + | – |
| COX2 | Protein-coding gene | 7,909 | 8,592 | 684 bp | + | 43.4% |
| tRNA-Lys | Transfer RNA | 8,596 | 8,663 | 68 bp | + | – |
| ATP8 | Protein-coding gene | 8,665 | 8,868 | 204 bp | + | 33.8% |
| ATP6 | Protein-coding gene | 8,826 | 9,506 | 681 bp | + | 44.2% |
| COX3 | Protein-coding gene | 9,506 | 10,289 | 784 bp | + | 46.8% |
| tRNA-Gly | Transfer RNA | 10,290 | 10,358 | 69 bp | + | – |
| ND3 | Protein-coding gene | 10,359 | 10,705 | 347 bp | + | 43.8% |
| tRNA-Arg | Transfer RNA | 10,706 | 10,772 | 67 bp | + | – |
| ND4L | Protein-coding gene | 10,773 | 11,069 | 297 bp | + | 39.7% |
| ND4 | Protein-coding gene | 11,063 | 12,440 | 1,378 bp | + | 42.0% |
| tRNA-His | Transfer RNA | 12,441 | 12,510 | 70 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 12,511 | 12,569 | 59 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 12,570 | 12,639 | 70 bp | + | – |
| ND5 | Protein-coding gene | 12,640 | 14,460 | 1,821 bp | + | 42.0% |
| ND6 | Protein-coding gene | 14,444 | 14,971 | 528 bp | − | 39.8% |
| tRNA-Glu | Transfer RNA | 14,972 | 15,040 | 69 bp | − | – |
| CYTB | Protein-coding gene | 15,044 | 16,183 | 1,140 bp | + | 44.0% |
| tRNA-Thr | Transfer RNA | 16,184 | 16,251 | 68 bp | + | – |
| tRNA-Pro | Transfer RNA | 16,252 | 16,317 | 66 bp | − | – |
| D-loop | Control region | 16,318 | 878 | 1,758 bp | – | 42.8% |