Phalacrocorax carbo
Great cormorant
GC content
45.21%
Whole genome
Non-coding
2,843 bp
D-loop 1,095 bp
Where Great cormorant sits among 2,335 species
Whole-genome GC content against maximum lifespan. Great cormorant has the 384th longest lifespan of them.
- This species
- Other birds
- Other classes
Base composition
Whole genome18,995 bp
- GC skew (G − C) / (G + C)
- −0.412
- AT skew (A − T) / (A + T)
- +0.161
Life history
- Maximum lifespan
- 32.1 years
- Adult weight
- 3.6 kg
- Birth weight
- 48.5 g
- Female sexual maturity
- 4 years
- Male sexual maturity
- 4 years
- Gestation or incubation
- 30 days
- Litter or clutch size
- 4
- Litters per year
- 1
- Postnatal growth rate
- 0.115 per day
Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 01715)
About this record
- Lifespan source
- AnAge build 15
- Based on
- an animal in the wild
- Data quality
- acceptable, large sample
- Licence
- Free to use, subject to the Human Ageing Genomic Resources conditions of use
- Status
- Updated in build 2
- In the database since
- build 1 (2015-07-01)
- Last changed
- build 2 (2026-11-13)
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome18,995 bp
Protein-coding11,903 bp
12S rRNA983 bp
16S rRNA1,517 bp
All tRNA1,760 bp
D-loop1,095 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 978 bp | 45.60% | −0.453 | +0.034 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,040 bp | 43.27% | −0.578 | +0.200 | |||||
| COX1Cytochrome c oxidase subunit I | 1,551 bp | 47.45% | −0.323 | +0.065 | |||||
| COX2Cytochrome c oxidase subunit II | 684 bp | 45.47% | −0.402 | +0.180 | |||||
| ATP8ATP synthase F0 subunit 8 | 168 bp | 40.48% | −0.794 | +0.280 | |||||
| ATP6ATP synthase F0 subunit 6 | 684 bp | 46.64% | −0.574 | +0.167 | |||||
| COX3Cytochrome c oxidase subunit III | 784 bp | 48.09% | −0.385 | +0.081 | |||||
| ND3NADH dehydrogenase subunit 3 | 352 bp | 43.18% | −0.513 | +0.090 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 46.80% | −0.482 | +0.139 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,378 bp | 45.86% | −0.566 | +0.169 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,818 bp | 45.82% | −0.539 | +0.200 | |||||
| ND6NADH dehydrogenase subunit 6 | 522 bp | 48.47% | −0.621 | +0.621 | |||||
| CYTBCytochrome b | 1,143 bp | 45.93% | −0.497 | +0.107 | |||||
| 12S rRNA12S ribosomal RNA | 983 bp | 51.98% | −0.217 | +0.267 | |||||
| 16S rRNA16S ribosomal RNA | 1,517 bp | 46.80% | −0.186 | +0.279 | |||||
| D-loopControl region (D-loop) | 1,095 bp | 35.43% | −0.196 | +0.030 |
Codon usage
Counts per codon, darker is more frequent3,806 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F5.5%
UUU47
UUC164
Leu L17.7%
UUA68
UUG18
CUU57
CUC155
CUA352
CUG23
Ile I7.5%
AUU57
AUC229
Met M4.4%
AUA143
AUG26
Val V4.1%
GUU28
GUC33
GUA77
GUG19
Ser S7.5%
UCU40
UCC87
UCA101
UCG4
AGU8
AGC47
Pro P5.9%
CCU19
CCC89
CCA111
CCG6
Thr T9.7%
ACU64
ACC151
ACA151
ACG5
Ala A7.9%
GCU62
GCC133
GCA99
GCG5
Tyr Y2.9%
UAU27
UAC82
His H2.9%
CAU22
CAC89
Gln Q2.6%
CAA93
CAG6
Asn N3.4%
AAU16
AAC112
Lys K2.3%
AAA84
AAG5
Asp D1.7%
GAU17
GAC49
Glu E2.4%
GAA82
GAG8
Cys C0.7%
UGU6
UGC21
Trp W2.8%
UGA98
UGG9
Arg R1.8%
CGU2
CGC18
CGA46
CGG4
Gly G5.8%
GGU23
GGC77
GGA90
GGG32
Stop 0.3%
UAA6
UAG1
AGA1
AGG2
Genome map
42 annotated features in NC_027267.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding14
- rRNA2
- tRNA25
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (45.2%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 42 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 70 | 70 bp | + | – |
| 12S rRNA | Ribosomal RNA | 71 | 1,053 | 983 bp | + | 52.0% |
| tRNA-Val | Transfer RNA | 1,054 | 1,124 | 71 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,165 | 2,681 | 1,517 bp | + | 46.8% |
| tRNA-Leu (UUR) | Transfer RNA | 2,729 | 2,802 | 74 bp | + | – |
| ND1 | Protein-coding gene | 2,808 | 3,785 | 978 bp | + | 45.6% |
| tRNA-Ile | Transfer RNA | 3,784 | 3,854 | 71 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,860 | 3,930 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 3,930 | 3,998 | 69 bp | + | – |
| ND2 | Protein-coding gene | 3,999 | 5,038 | 1,040 bp | + | 43.3% |
| tRNA-Trp | Transfer RNA | 5,039 | 5,108 | 70 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,110 | 5,178 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,185 | 5,257 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,261 | 5,327 | 67 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,327 | 5,397 | 71 bp | − | – |
| COX1 | Protein-coding gene | 5,399 | 6,949 | 1,551 bp | + | 47.5% |
| tRNA-Ser (UCN) | Transfer RNA | 6,941 | 7,015 | 75 bp | − | – |
| tRNA-Asp | Transfer RNA | 7,018 | 7,086 | 69 bp | + | – |
| COX2 | Protein-coding gene | 7,088 | 7,771 | 684 bp | + | 45.5% |
| tRNA-Lys | Transfer RNA | 7,773 | 7,845 | 73 bp | + | – |
| ATP8 | Protein-coding gene | 7,846 | 8,013 | 168 bp | + | 40.5% |
| ATP6 | Protein-coding gene | 8,004 | 8,687 | 684 bp | + | 46.6% |
| COX3 | Protein-coding gene | 8,687 | 9,470 | 784 bp | + | 48.1% |
| tRNA-Gly | Transfer RNA | 9,471 | 9,539 | 69 bp | + | – |
| ND3 | Protein-coding gene | 9,540 | 9,891 | 352 bp | + | 43.2% |
| tRNA-Arg | Transfer RNA | 9,894 | 9,962 | 69 bp | + | – |
| ND4L | Protein-coding gene | 9,964 | 10,260 | 297 bp | + | 46.8% |
| ND4 | Protein-coding gene | 10,254 | 11,631 | 1,378 bp | + | 45.9% |
| tRNA-His | Transfer RNA | 11,632 | 11,701 | 70 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,702 | 11,769 | 68 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,770 | 11,840 | 71 bp | + | – |
| ND5 | Protein-coding gene | 11,841 | 13,658 | 1,818 bp | + | 45.8% |
| CYTB | Protein-coding gene | 13,670 | 14,812 | 1,143 bp | + | 45.9% |
| tRNA-Thr | Transfer RNA | 14,817 | 14,886 | 70 bp | + | – |
| tRNA-Pro | Transfer RNA | 14,896 | 14,965 | 70 bp | − | – |
| ND6 | Protein-coding gene | 14,974 | 15,495 | 522 bp | − | 48.5% |
| tRNA-Glu | Transfer RNA | 15,499 | 15,569 | 71 bp | − | – |
| tRNA-Thr | Transfer RNA | 17,148 | 17,217 | 70 bp | + | – |
| tRNA-Pro | Transfer RNA | 17,227 | 17,296 | 70 bp | − | – |
| ND6 | Protein-coding gene | 17,305 | 17,826 | 522 bp | − | 48.5% |
| tRNA-Glu | Transfer RNA | 17,830 | 17,900 | 71 bp | − | – |
| D-loop | Control region | 17,901 | 18,995 | 1,095 bp | – | 35.4% |