Romanogobio ciscaucasicus

North Caucasian longbarbel gudgeon

Maximum lifespan

6 yr

FishBase record

mtDNA size

16,603 bp

RefSeq NC_031558.1

GC content

43.55%

Whole genome

Non-coding

988 bp

D-loop 928 bp

Where North Caucasian longbarbel gudgeon sits among 2,335 species

Whole-genome GC content against maximum lifespan. North Caucasian longbarbel gudgeon has the 1,943rd longest lifespan of them.

This species
Other ray-finned fishes
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome16,603 bp

GC skew (G − C) / (G + C): −0.233
AT skew (A − T) / (A + T): +0.068

Life history

Maximum lifespan: 6 years

Lifespan from FishBase · FishBase record

About this record

Lifespan source: FishBase v26.06
Based on: an animal in the wild
Data quality: acceptable
Licence: CC BY-NC 4.0
Status: New in build 2
In the database since: build 2 (2026-11-13)

The lifespan value is for non-commercial use only; the genome data is not restricted by it.

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome16,603 bp

Protein-coding11,403 bp

12S rRNA959 bp

16S rRNA1,694 bp

All tRNA1,559 bp

D-loop928 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Romanogobio ciscaucasicus genes

ND1NADH dehydrogenase subunit 1	975 bp	28.8%	27.7%	15.6%	27.9%	43.49%	−0.283	+0.020
ND2NADH dehydrogenase subunit 2	1,045 bp	30.1%	24.3%	14.0%	31.6%	45.55%	−0.387	+0.107
COX1Cytochrome c oxidase subunit I	1,551 bp	25.1%	30.0%	18.0%	26.9%	44.87%	−0.198	−0.088
COX2Cytochrome c oxidase subunit II	691 bp	30.7%	26.9%	15.9%	26.5%	42.40%	−0.249	+0.065
ATP8ATP synthase F0 subunit 8	165 bp	36.4%	24.2%	10.9%	28.5%	39.39%	−0.446	+0.200
ATP6ATP synthase F0 subunit 6	683 bp	27.2%	30.6%	14.2%	28.0%	42.17%	−0.326	−0.058
COX3Cytochrome c oxidase subunit III	784 bp	27.4%	27.8%	17.2%	27.6%	44.77%	−0.231	−0.007
ND3NADH dehydrogenase subunit 3	349 bp	26.9%	28.9%	15.2%	28.9%	44.13%	−0.312	−0.036
ND4LNADH dehydrogenase subunit 4L	297 bp	25.3%	26.6%	16.2%	32.0%	48.15%	−0.329	−0.026
ND4NADH dehydrogenase subunit 4	1,382 bp	28.4%	27.1%	15.3%	29.2%	44.50%	−0.314	+0.025
ND5NADH dehydrogenase subunit 5	1,836 bp	31.6%	27.3%	13.9%	27.1%	41.01%	−0.320	+0.073
ND6NADH dehydrogenase subunit 6	522 bp	38.7%	16.5%	14.2%	30.7%	44.83%	−0.368	+0.403
CYTBCytochrome b	1,141 bp	27.6%	28.7%	15.4%	28.2%	43.65%	−0.293	−0.020
12S rRNA12S ribosomal RNA	959 bp	31.3%	19.1%	22.3%	27.3%	49.64%	−0.101	+0.242
16S rRNA16S ribosomal RNA	1,694 bp	35.3%	20.7%	20.8%	23.1%	43.98%	−0.052	+0.260
D-loopControl region (D-loop)	928 bp	33.1%	33.5%	13.5%	19.9%	33.41%	−0.194	−0.006

Codon usage

Counts per codon, darker is more frequent

Gene

3,804 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F6.0%

UUU109

UUC119

Leu L16.3%

UUA126

UUG19

CUU117

CUC95

CUA211

CUG52

Ile I7.9%

AUU189

AUC110

Met M4.7%

AUA122

AUG56

Val V5.7%

GUU40

GUC51

GUA99

GUG28

Ser S6.3%

UCU40

UCC54

UCA86

UCG5

AGU13

AGC43

Pro P5.7%

CCU40

CCC71

CCA90

CCG16

Thr T8.1%

ACU51

ACC94

ACA146

ACG16

Ala A8.6%

GCU58

GCC148

GCA105

GCG18

Tyr Y2.9%

UAU52

UAC59

His H2.7%

CAU40

CAC63

Gln Q2.5%

CAA76

CAG21

Asn N3.3%

AAU41

AAC85

Lys K2.1%

AAA69

AAG9

Asp D2.1%

GAU21

GAC59

Glu E2.6%

GAA73

GAG27

Cys C0.7%

UGU14

UGC12

Trp W3.2%

UGA103

UGG18

Arg R2.0%

CGU8

CGC15

CGA44

CGG10

Gly G6.4%

GGU23

GGC64

GGA103

GGG52

Stop 0.2%

UAA3

UAG3

AGA0

AGG0

Genome map

38 annotated features in NC_031558.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA22
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (43.6%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 38 features as a table

Annotated features of Romanogobio ciscaucasicus (NC_031558.1)
Feature	Type	Start	End	Length	Strand	GC
tRNA-Phe	Transfer RNA	1	69	69 bp	+	–
12S rRNA	Ribosomal RNA	70	1,028	959 bp	+	49.6%
tRNA-Val	Transfer RNA	1,029	1,100	72 bp	+	–
16S rRNA	Ribosomal RNA	1,101	2,794	1,694 bp	+	44.0%
tRNA-Leu (UUR)	Transfer RNA	2,795	2,870	76 bp	+	–
ND1	Protein-coding gene	2,872	3,846	975 bp	+	43.5%
tRNA-Ile	Transfer RNA	3,851	3,922	72 bp	+	–
tRNA-Gln	Transfer RNA	3,921	3,991	71 bp	−	–
tRNA-Met	Transfer RNA	3,993	4,061	69 bp	+	–
ND2	Protein-coding gene	4,062	5,106	1,045 bp	+	45.6%
tRNA-Trp	Transfer RNA	5,107	5,177	71 bp	+	–
tRNA-Ala	Transfer RNA	5,179	5,247	69 bp	−	–
tRNA-Asn	Transfer RNA	5,249	5,321	73 bp	−	–
tRNA-Cys	Transfer RNA	5,352	5,419	68 bp	−	–
tRNA-Tyr	Transfer RNA	5,421	5,491	71 bp	−	–
COX1	Protein-coding gene	5,493	7,043	1,551 bp	+	44.9%
tRNA-Ser (UCN)	Transfer RNA	7,044	7,114	71 bp	−	–
tRNA-Asp	Transfer RNA	7,118	7,189	72 bp	+	–
COX2	Protein-coding gene	7,203	7,893	691 bp	+	42.4%
tRNA-Lys	Transfer RNA	7,894	7,969	76 bp	+	–
ATP8	Protein-coding gene	7,971	8,135	165 bp	+	39.4%
ATP6	Protein-coding gene	8,129	8,811	683 bp	+	42.2%
COX3	Protein-coding gene	8,812	9,595	784 bp	+	44.8%
tRNA-Gly	Transfer RNA	9,596	9,666	71 bp	+	–
ND3	Protein-coding gene	9,667	10,015	349 bp	+	44.1%
tRNA-Arg	Transfer RNA	10,016	10,085	70 bp	+	–
ND4L	Protein-coding gene	10,086	10,382	297 bp	+	48.2%
ND4	Protein-coding gene	10,376	11,757	1,382 bp	+	44.5%
tRNA-His	Transfer RNA	11,758	11,826	69 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	11,827	11,895	69 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	11,897	11,969	73 bp	+	–
ND5	Protein-coding gene	11,970	13,805	1,836 bp	+	41.0%
ND6	Protein-coding gene	13,802	14,323	522 bp	−	44.8%
tRNA-Glu	Transfer RNA	14,324	14,392	69 bp	−	–
CYTB	Protein-coding gene	14,395	15,535	1,141 bp	+	43.7%
tRNA-Thr	Transfer RNA	15,536	15,607	72 bp	+	–
tRNA-Pro	Transfer RNA	15,606	15,675	70 bp	−	–
D-loop	Control region	15,676	16,603	928 bp	–	33.4%