Smaug warreni

Warren's spinytail lizard

Maximum lifespan

19.8 yr

AnAge record

mtDNA size

17,184 bp

RefSeq NC_005962.1

GC content

44.23%

Whole genome

Non-coding

1,766 bp

D-loop 1,588 bp

Where Warren's spinytail lizard sits among 2,335 species

Whole-genome GC content against maximum lifespan. Warren's spinytail lizard has the 959th longest lifespan of them.

This species
Other reptiles
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome17,184 bp

GC skew (G − C) / (G + C): −0.356
AT skew (A − T) / (A + T): +0.124

Life history

Maximum lifespan: 19.8 years

Lifespan from AnAge · AnAge record

About this record

Lifespan source: AnAge build 15
Based on: an animal in captivity
Data quality: acceptable, medium sample
Licence: Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status: New in build 2
In the database since: build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome17,184 bp

Protein-coding11,354 bp

12S rRNA896 bp

16S rRNA1,509 bp

All tRNA1,665 bp

D-loop1,588 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Smaug warreni genes

ND1NADH dehydrogenase subunit 1	965 bp	30.3%	23.9%	13.3%	32.5%	45.80%	−0.421	+0.117
ND2NADH dehydrogenase subunit 2	1,027 bp	34.0%	22.9%	10.5%	32.6%	43.14%	−0.512	+0.195
COX1Cytochrome c oxidase subunit I	1,560 bp	27.5%	27.6%	17.2%	27.7%	44.87%	−0.234	−0.002
COX2Cytochrome c oxidase subunit II	688 bp	31.0%	25.6%	14.2%	29.2%	43.46%	−0.344	+0.095
ATP8ATP synthase F0 subunit 8	165 bp	33.9%	24.2%	10.9%	30.9%	41.82%	−0.478	+0.167
ATP6ATP synthase F0 subunit 6	683 bp	30.0%	26.8%	9.8%	33.4%	43.19%	−0.546	+0.057
COX3Cytochrome c oxidase subunit III	784 bp	24.5%	26.9%	16.8%	31.8%	48.60%	−0.307	−0.047
ND3NADH dehydrogenase subunit 3	346 bp	28.0%	28.0%	11.6%	32.4%	43.93%	−0.474	0.000
ND4LNADH dehydrogenase subunit 4L	297 bp	26.9%	26.6%	13.5%	33.0%	46.47%	−0.420	+0.006
ND4NADH dehydrogenase subunit 4	1,374 bp	29.0%	26.3%	10.8%	33.9%	44.69%	−0.518	+0.050
ND5NADH dehydrogenase subunit 5	1,824 bp	32.5%	24.1%	10.9%	32.6%	43.48%	−0.498	+0.148
ND6NADH dehydrogenase subunit 6	519 bp	43.4%	12.5%	10.6%	33.5%	44.12%	−0.520	+0.552
CYTBCytochrome b	1,143 bp	28.2%	27.2%	11.6%	33.0%	44.62%	−0.478	+0.017
12S rRNA12S ribosomal RNA	896 bp	34.4%	21.7%	18.9%	25.1%	43.97%	−0.142	+0.227
16S rRNA16S ribosomal RNA	1,509 bp	35.9%	20.0%	18.2%	25.9%	44.07%	−0.176	+0.284
D-loopControl region (D-loop)	1,588 bp	30.2%	25.3%	17.9%	26.7%	44.58%	−0.198	+0.089

Codon usage

Counts per codon, darker is more frequent

Gene

3,789 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F6.0%

UUU113

UUC113

Leu L16.9%

UUA78

UUG21

CUU127

CUC146

CUA228

CUG41

Ile I7.8%

AUU141

AUC153

Met M5.3%

AUA168

AUG32

Val V4.5%

GUU47

GUC48

GUA59

GUG18

Ser S6.7%

UCU39

UCC75

UCA92

UCG7

AGU8

AGC32

Pro P5.8%

CCU22

CCC64

CCA112

CCG21

Thr T10.1%

ACU30

ACC169

ACA171

ACG11

Ala A7.2%

GCU32

GCC141

GCA93

GCG5

Tyr Y2.7%

UAU33

UAC70

His H2.8%

CAU17

CAC89

Gln Q2.5%

CAA88

CAG8

Asn N3.6%

AAU25

AAC113

Lys K2.4%

AAA83

AAG9

Asp D1.7%

GAU16

GAC49

Glu E2.5%

GAA67

GAG28

Cys C0.9%

UGU7

UGC27

Trp W2.8%

UGA89

UGG17

Arg R1.9%

CGU10

CGC23

CGA37

CGG2

Gly G5.8%

GGU26

GGC81

GGA72

GGG39

Stop 0.2%

UAA4

UAG2

AGA1

AGG0

Genome map

40 annotated features in NC_005962.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA24
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (44.2%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 40 features as a table

Annotated features of Smaug warreni (NC_005962.1)
Feature	Type	Start	End	Length	Strand	GC
tRNA-Phe	Transfer RNA	1	72	72 bp	+	–
12S rRNA	Ribosomal RNA	73	968	896 bp	+	44.0%
tRNA-Val	Transfer RNA	968	1,038	71 bp	+	–
16S rRNA	Ribosomal RNA	1,039	2,547	1,509 bp	+	44.1%
tRNA-Leu (UUR)	Transfer RNA	2,548	2,624	77 bp	+	–
ND1	Protein-coding gene	2,625	3,589	965 bp	+	45.8%
tRNA-Ile	Transfer RNA	3,590	3,659	70 bp	+	–
tRNA-Gln	Transfer RNA	3,658	3,728	71 bp	−	–
tRNA-Met	Transfer RNA	3,728	3,796	69 bp	+	–
ND2	Protein-coding gene	3,797	4,823	1,027 bp	+	43.1%
tRNA-Trp	Transfer RNA	4,824	4,896	73 bp	+	–
tRNA-Ala	Transfer RNA	4,898	4,966	69 bp	−	–
tRNA-Asn	Transfer RNA	4,968	5,040	73 bp	−	–
tRNA-Cys	Transfer RNA	5,068	5,130	63 bp	−	–
tRNA-Tyr	Transfer RNA	5,131	5,197	67 bp	−	–
COX1	Protein-coding gene	5,199	6,758	1,560 bp	+	44.9%
tRNA-Ser (UCN)	Transfer RNA	6,754	6,824	71 bp	−	–
tRNA-Asp	Transfer RNA	6,828	6,895	68 bp	+	–
COX2	Protein-coding gene	6,897	7,584	688 bp	+	43.5%
tRNA-Lys	Transfer RNA	7,585	7,650	66 bp	+	–
ATP8	Protein-coding gene	7,652	7,816	165 bp	+	41.8%
ATP6	Protein-coding gene	7,807	8,489	683 bp	+	43.2%
COX3	Protein-coding gene	8,490	9,273	784 bp	+	48.6%
tRNA-Gly	Transfer RNA	9,274	9,342	69 bp	+	–
ND3	Protein-coding gene	9,343	9,688	346 bp	+	43.9%
tRNA-Arg	Transfer RNA	9,689	9,757	69 bp	+	–
ND4L	Protein-coding gene	9,758	10,054	297 bp	+	46.5%
ND4	Protein-coding gene	10,048	11,421	1,374 bp	+	44.7%
tRNA-His	Transfer RNA	11,422	11,490	69 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	11,491	11,556	66 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	11,556	11,626	71 bp	+	–
ND5	Protein-coding gene	11,628	13,451	1,824 bp	+	43.5%
ND6	Protein-coding gene	13,448	13,966	519 bp	−	44.1%
tRNA-Glu	Transfer RNA	13,967	14,035	69 bp	−	–
CYTB	Protein-coding gene	14,040	15,182	1,143 bp	+	44.6%
tRNA-Thr	Transfer RNA	15,189	15,257	69 bp	+	–
tRNA-Pro	Transfer RNA	15,258	15,326	69 bp	−	–
tRNA-Thr	Transfer RNA	15,459	15,527	69 bp	+	–
tRNA-Pro	Transfer RNA	15,528	15,596	69 bp	−	–
D-loop	Control region	15,597	17,184	1,588 bp	–	44.6%