Testudo graeca
Mediterranean spur-thighed tortoise
GC content
37.29%
Whole genome
Non-coding
3,820 bp
D-loop 3,771 bp
Where Mediterranean spur-thighed tortoise sits among 2,335 species
Whole-genome GC content against maximum lifespan. Mediterranean spur-thighed tortoise has the 13th longest lifespan of them.
- This species
- Other reptiles
- Other classes
Base composition
Whole genome19,278 bp
- GC skew (G − C) / (G + C)
- −0.368
- AT skew (A − T) / (A + T)
- +0.087
Life history
- Maximum lifespan
- 127 years
- Adult weight
- 1.4 kg
- Female sexual maturity
- 12.5 years
- Male sexual maturity
- 12.5 years
Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 03782)
About this record
- Lifespan source
- AnAge build 15
- Based on
- an animal in captivity
- Data quality
- acceptable, large sample
- Licence
- Free to use, subject to the Human Ageing Genomic Resources conditions of use
- Status
- Updated in build 2
- In the database since
- build 1 (2015-07-01)
- Last changed
- build 2 (2026-11-13)
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome19,278 bp
Protein-coding11,359 bp
12S rRNA970 bp
16S rRNA1,600 bp
All tRNA1,542 bp
D-loop3,771 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 972 bp | 40.95% | −0.447 | +0.094 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,041 bp | 39.29% | −0.545 | +0.196 | |||||
| COX1Cytochrome c oxidase subunit I | 1,548 bp | 40.96% | −0.221 | +0.002 | |||||
| COX2Cytochrome c oxidase subunit II | 687 bp | 37.99% | −0.287 | +0.131 | |||||
| ATP8ATP synthase F0 subunit 8 | 174 bp | 37.36% | −0.692 | +0.339 | |||||
| ATP6ATP synthase F0 subunit 6 | 684 bp | 37.14% | −0.504 | +0.121 | |||||
| COX3Cytochrome c oxidase subunit III | 784 bp | 42.22% | −0.263 | +0.064 | |||||
| ND3NADH dehydrogenase subunit 3 | 351 bp | 39.03% | −0.314 | 0.000 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 41.41% | −0.447 | −0.023 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,378 bp | 35.85% | −0.462 | +0.131 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,797 bp | 38.56% | −0.483 | +0.154 | |||||
| ND6NADH dehydrogenase subunit 6 | 525 bp | 37.52% | −0.604 | +0.537 | |||||
| CYTBCytochrome b | 1,144 bp | 42.05% | −0.455 | +0.083 | |||||
| 12S rRNA12S ribosomal RNA | 970 bp | 40.00% | −0.191 | +0.247 | |||||
| 16S rRNA16S ribosomal RNA | 1,600 bp | 38.31% | −0.139 | +0.291 | |||||
| D-loopControl region (D-loop) | 3,771 bp | 29.09% | −0.437 | −0.139 |
Codon usage
Counts per codon, darker is more frequent3,793 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F5.7%
UUU97
UUC120
Leu L16.0%
UUA145
UUG21
CUU58
CUC61
CUA300
CUG21
Ile I8.3%
AUU150
AUC163
Met M6.5%
AUA213
AUG33
Val V4.2%
GUU37
GUC21
GUA91
GUG11
Ser S7.3%
UCU42
UCC67
UCA119
UCG3
AGU9
AGC37
Pro P5.5%
CCU23
CCC34
CCA143
CCG8
Thr T9.4%
ACU76
ACC124
ACA154
ACG4
Ala A6.4%
GCU51
GCC103
GCA85
GCG3
Tyr Y3.3%
UAU44
UAC82
His H2.7%
CAU41
CAC62
Gln Q2.6%
CAA94
CAG6
Asn N4.1%
AAU49
AAC106
Lys K2.4%
AAA83
AAG8
Asp D1.6%
GAU28
GAC33
Glu E2.4%
GAA77
GAG13
Cys C0.8%
UGU16
UGC14
Trp W2.9%
UGA102
UGG8
Arg R1.8%
CGU7
CGC13
CGA47
CGG3
Gly G5.8%
GGU34
GGC57
GGA106
GGG23
Stop 0.3%
UAA6
UAG2
AGA0
AGG2
Genome map
38 annotated features in NC_007692.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (37.3%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 69 | 69 bp | + | – |
| 12S rRNA | Ribosomal RNA | 70 | 1,039 | 970 bp | + | 40.0% |
| tRNA-Val | Transfer RNA | 1,040 | 1,109 | 70 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,110 | 2,709 | 1,600 bp | + | 38.3% |
| tRNA-Leu (UUR) | Transfer RNA | 2,710 | 2,785 | 76 bp | + | – |
| ND1 | Protein-coding gene | 2,786 | 3,757 | 972 bp | + | 41.0% |
| tRNA-Ile | Transfer RNA | 3,757 | 3,826 | 70 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,826 | 3,896 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 3,896 | 3,964 | 69 bp | + | – |
| ND2 | Protein-coding gene | 3,969 | 5,009 | 1,041 bp | + | 39.3% |
| tRNA-Trp | Transfer RNA | 5,008 | 5,085 | 78 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,087 | 5,155 | 69 bp | + | – |
| tRNA-Asn | Transfer RNA | 5,158 | 5,230 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,258 | 5,323 | 66 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,324 | 5,394 | 71 bp | − | – |
| COX1 | Protein-coding gene | 5,396 | 6,943 | 1,548 bp | + | 41.0% |
| tRNA-Ser (UCN) | Transfer RNA | 6,935 | 7,005 | 71 bp | + | – |
| tRNA-Asp | Transfer RNA | 7,006 | 7,075 | 70 bp | + | – |
| COX2 | Protein-coding gene | 7,076 | 7,762 | 687 bp | + | 38.0% |
| tRNA-Lys | Transfer RNA | 7,768 | 7,840 | 73 bp | + | – |
| ATP8 | Protein-coding gene | 7,842 | 8,015 | 174 bp | + | 37.4% |
| ATP6 | Protein-coding gene | 8,006 | 8,689 | 684 bp | + | 37.1% |
| COX3 | Protein-coding gene | 8,689 | 9,472 | 784 bp | + | 42.2% |
| tRNA-Gly | Transfer RNA | 9,473 | 9,541 | 69 bp | + | – |
| ND3 | Protein-coding gene | 9,542 | 9,893 | 352 bp | + | 39.0% |
| tRNA-Arg | Transfer RNA | 9,893 | 9,961 | 69 bp | + | – |
| ND4L | Protein-coding gene | 9,962 | 10,258 | 297 bp | + | 41.4% |
| ND4 | Protein-coding gene | 10,252 | 11,629 | 1,378 bp | + | 35.9% |
| tRNA-His | Transfer RNA | 11,630 | 11,697 | 68 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,698 | 11,763 | 66 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,763 | 11,833 | 71 bp | + | – |
| ND5 | Protein-coding gene | 11,836 | 13,632 | 1,797 bp | + | 38.6% |
| ND6 | Protein-coding gene | 13,628 | 14,152 | 525 bp | − | 37.5% |
| tRNA-Glu | Transfer RNA | 14,153 | 14,219 | 67 bp | − | – |
| CYTB | Protein-coding gene | 14,224 | 15,367 | 1,144 bp | + | 42.1% |
| tRNA-Thr | Transfer RNA | 15,368 | 15,437 | 70 bp | + | – |
| tRNA-Pro | Transfer RNA | 15,439 | 15,507 | 69 bp | + | – |
| D-loop | Control region | 15,508 | 19,278 | 3,771 bp | – | 29.1% |