Testudo kleinmanni

Egyptian tortoise

Maximum lifespan

21 yr

AnAge record

mtDNA size

17,670 bp

RefSeq NC_007699.1

GC content

38.85%

Whole genome

Non-coding

2,191 bp

D-loop 2,151 bp

Where Egyptian tortoise sits among 2,335 species

Whole-genome GC content against maximum lifespan. Egyptian tortoise has the 845th longest lifespan of them.

This species
Other reptiles
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome17,670 bp

GC skew (G − C) / (G + C): −0.354
AT skew (A − T) / (A + T): +0.116

Life history

Maximum lifespan: 21 years
Adult weight: 440 g
Female sexual maturity: 6 years

Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 03784)

About this record

Lifespan source: AnAge build 15
Based on: an animal in captivity
Data quality: acceptable, large sample
Licence: Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status: Updated in build 2
In the database since: build 1 (2015-07-01)
Last changed: build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome17,670 bp

Protein-coding11,368 bp

12S rRNA968 bp

16S rRNA1,603 bp

All tRNA1,553 bp

D-loop2,151 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Testudo kleinmanni genes

ND1NADH dehydrogenase subunit 1	972 bp	32.2%	27.0%	11.2%	29.6%	40.84%	−0.451	+0.089
ND2NADH dehydrogenase subunit 2	1,041 bp	35.9%	25.1%	9.2%	29.8%	39.00%	−0.527	+0.178
COX1Cytochrome c oxidase subunit I	1,548 bp	29.2%	29.7%	16.6%	24.5%	41.15%	−0.193	−0.008
COX2Cytochrome c oxidase subunit II	687 bp	35.1%	26.5%	12.7%	25.8%	38.43%	−0.341	+0.139
ATP8ATP synthase F0 subunit 8	183 bp	43.2%	19.1%	6.0%	31.7%	37.71%	−0.681	+0.386
ATP6ATP synthase F0 subunit 6	684 bp	34.9%	27.6%	9.2%	28.2%	37.43%	−0.508	+0.117
COX3Cytochrome c oxidase subunit III	784 bp	31.4%	26.9%	14.8%	26.9%	41.71%	−0.291	+0.077
ND3NADH dehydrogenase subunit 3	351 bp	31.3%	30.8%	12.8%	25.1%	37.89%	−0.323	+0.009
ND4LNADH dehydrogenase subunit 4L	297 bp	29.3%	29.3%	11.4%	30.0%	41.41%	−0.447	0.000
ND4NADH dehydrogenase subunit 4	1,378 bp	35.1%	27.6%	10.6%	26.7%	37.30%	−0.432	+0.120
ND5NADH dehydrogenase subunit 5	1,797 bp	34.8%	25.5%	10.0%	29.6%	39.62%	−0.494	+0.154
ND6NADH dehydrogenase subunit 6	525 bp	47.4%	14.9%	9.0%	28.8%	37.71%	−0.525	+0.523
CYTBCytochrome b	1,144 bp	31.3%	26.1%	11.7%	30.9%	42.57%	−0.450	+0.090
12S rRNA12S ribosomal RNA	968 bp	36.8%	21.2%	16.6%	25.4%	42.05%	−0.209	+0.269
16S rRNA16S ribosomal RNA	1,603 bp	40.2%	20.6%	16.2%	23.1%	39.24%	−0.176	+0.322
D-loopControl region (D-loop)	2,151 bp	29.7%	37.9%	10.7%	21.7%	32.40%	−0.340	−0.121

Codon usage

Counts per codon, darker is more frequent

Gene

3,796 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F5.6%

UUU99

UUC114

Leu L16.2%

UUA125

UUG17

CUU80

CUC58

CUA304

CUG30

Ile I8.5%

AUU156

AUC168

Met M6.3%

AUA203

AUG38

Val V4.0%

GUU27

GUC25

GUA86

GUG13

Ser S7.4%

UCU46

UCC58

UCA127

UCG4

AGU12

AGC34

Pro P5.5%

CCU18

CCC40

CCA139

CCG11

Thr T9.2%

ACU75

ACC123

ACA147

ACG3

Ala A6.5%

GCU48

GCC106

GCA91

GCG1

Tyr Y3.3%

UAU50

UAC77

His H2.7%

CAU41

CAC61

Gln Q2.7%

CAA101

CAG2

Asn N4.1%

AAU36

AAC119

Lys K2.4%

AAA86

AAG4

Asp D1.7%

GAU29

GAC35

Glu E2.4%

GAA81

GAG9

Cys C0.8%

UGU17

UGC13

Trp W2.9%

UGA100

UGG11

Arg R1.8%

CGU5

CGC13

CGA51

CGG0

Gly G5.8%

GGU30

GGC59

GGA95

GGG35

Stop 0.3%

UAA6

UAG2

AGA0

AGG2

Genome map

38 annotated features in NC_007699.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA22
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (38.8%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 38 features as a table

Annotated features of Testudo kleinmanni (NC_007699.1)
Feature	Type	Start	End	Length	Strand	GC
tRNA-Phe	Transfer RNA	1	69	69 bp	+	–
12S rRNA	Ribosomal RNA	70	1,037	968 bp	+	42.1%
tRNA-Val	Transfer RNA	1,038	1,108	71 bp	+	–
16S rRNA	Ribosomal RNA	1,109	2,711	1,603 bp	+	39.2%
tRNA-Leu (UUR)	Transfer RNA	2,712	2,787	76 bp	+	–
ND1	Protein-coding gene	2,788	3,759	972 bp	+	40.8%
tRNA-Ile	Transfer RNA	3,759	3,828	70 bp	+	–
tRNA-Gln	Transfer RNA	3,829	3,899	71 bp	−	–
tRNA-Met	Transfer RNA	3,899	3,967	69 bp	+	–
ND2	Protein-coding gene	3,972	5,012	1,041 bp	+	39.0%
tRNA-Trp	Transfer RNA	5,011	5,088	78 bp	+	–
tRNA-Ala	Transfer RNA	5,090	5,158	69 bp	−	–
tRNA-Asn	Transfer RNA	5,161	5,234	74 bp	−	–
tRNA-Cys	Transfer RNA	5,260	5,325	66 bp	−	–
tRNA-Tyr	Transfer RNA	5,326	5,397	72 bp	−	–
COX1	Protein-coding gene	5,398	6,945	1,548 bp	+	41.2%
tRNA-Ser (UCN)	Transfer RNA	6,937	7,007	71 bp	+	–
tRNA-Asp	Transfer RNA	7,008	7,077	70 bp	+	–
COX2	Protein-coding gene	7,078	7,764	687 bp	+	38.4%
tRNA-Lys	Transfer RNA	7,765	7,842	78 bp	+	–
ATP8	Protein-coding gene	7,844	8,026	183 bp	+	37.7%
ATP6	Protein-coding gene	8,017	8,700	684 bp	+	37.4%
COX3	Protein-coding gene	8,700	9,483	784 bp	+	41.7%
tRNA-Gly	Transfer RNA	9,484	9,552	69 bp	+	–
ND3	Protein-coding gene	9,553	9,904	352 bp	+	37.9%
tRNA-Arg	Transfer RNA	9,904	9,972	69 bp	+	–
ND4L	Protein-coding gene	9,973	10,269	297 bp	+	41.4%
ND4	Protein-coding gene	10,263	11,640	1,378 bp	+	37.3%
tRNA-His	Transfer RNA	11,641	11,708	68 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	11,709	11,774	66 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	11,774	11,844	71 bp	+	–
ND5	Protein-coding gene	11,847	13,643	1,797 bp	+	39.6%
ND6	Protein-coding gene	13,639	14,163	525 bp	−	37.7%
tRNA-Glu	Transfer RNA	14,164	14,230	67 bp	+	–
CYTB	Protein-coding gene	14,235	15,378	1,144 bp	+	42.6%
tRNA-Thr	Transfer RNA	15,379	15,449	71 bp	+	–
tRNA-Pro	Transfer RNA	15,450	15,519	70 bp	+	–
D-loop	Control region	15,520	17,670	2,151 bp	–	32.4%