Testudo marginata

Marginated tortoise

Maximum lifespan

21.2 yr

AnAge record

mtDNA size

19,438 bp

RefSeq NC_007698.1

GC content

38.28%

Whole genome

Non-coding

3,993 bp

D-loop 3,943 bp

Where Marginated tortoise sits among 2,335 species

Whole-genome GC content against maximum lifespan. Marginated tortoise has the 829th longest lifespan of them.

This species
Other reptiles
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome19,438 bp

GC skew (G − C) / (G + C): −0.355
AT skew (A − T) / (A + T): +0.098

Life history

Maximum lifespan: 21.2 years
Adult weight: 1.9 kg

Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 03785)

About this record

Lifespan source: AnAge build 15
Based on: an animal in captivity
Data quality: acceptable, medium sample
Licence: Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status: Updated in build 2
In the database since: build 1 (2015-07-01)
Last changed: build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome19,438 bp

Protein-coding11,353 bp

12S rRNA968 bp

16S rRNA1,597 bp

All tRNA1,540 bp

D-loop3,943 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Testudo marginata genes

ND1NADH dehydrogenase subunit 1	972 bp	32.8%	26.7%	10.9%	29.5%	40.43%	−0.461	+0.102
ND2NADH dehydrogenase subunit 2	1,041 bp	36.7%	23.9%	8.6%	30.7%	39.39%	−0.561	+0.211
COX1Cytochrome c oxidase subunit I	1,548 bp	29.4%	29.3%	16.6%	24.7%	41.34%	−0.197	+0.002
COX2Cytochrome c oxidase subunit II	687 bp	34.8%	26.3%	13.4%	25.5%	38.87%	−0.311	+0.138
ATP8ATP synthase F0 subunit 8	174 bp	40.8%	20.7%	6.9%	31.6%	38.51%	−0.642	+0.327
ATP6ATP synthase F0 subunit 6	684 bp	35.5%	28.2%	8.8%	27.5%	36.26%	−0.516	+0.115
COX3Cytochrome c oxidase subunit III	784 bp	31.1%	27.2%	15.1%	26.7%	41.71%	−0.278	+0.068
ND3NADH dehydrogenase subunit 3	351 bp	29.9%	30.8%	13.4%	25.9%	39.32%	−0.319	−0.014
ND4LNADH dehydrogenase subunit 4L	297 bp	29.6%	29.6%	11.4%	29.3%	40.74%	−0.438	0.000
ND4NADH dehydrogenase subunit 4	1,378 bp	35.1%	26.5%	10.7%	27.7%	38.46%	−0.442	+0.139
ND5NADH dehydrogenase subunit 5	1,791 bp	34.7%	25.6%	10.4%	29.3%	39.70%	−0.477	+0.152
ND6NADH dehydrogenase subunit 6	525 bp	48.0%	13.0%	8.2%	30.9%	39.05%	−0.580	+0.575
CYTBCytochrome b	1,144 bp	31.7%	26.7%	11.3%	30.3%	41.61%	−0.458	+0.087
12S rRNA12S ribosomal RNA	968 bp	36.8%	21.9%	16.8%	24.5%	41.32%	−0.185	+0.254
16S rRNA16S ribosomal RNA	1,597 bp	40.3%	21.0%	16.0%	22.7%	38.70%	−0.175	+0.316
D-loopControl region (D-loop)	3,943 bp	30.3%	37.4%	10.5%	21.8%	32.29%	−0.351	−0.106

Codon usage

Counts per codon, darker is more frequent

Gene

3,791 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F5.8%

UUU91

UUC127

Leu L16.0%

UUA122

UUG23

CUU69

CUC58

CUA308

CUG27

Ile I8.4%

AUU147

AUC172

Met M6.6%

AUA215

AUG34

Val V3.9%

GUU31

GUC23

GUA83

GUG11

Ser S7.3%

UCU47

UCC56

UCA120

UCG8

AGU10

AGC35

Pro P5.5%

CCU21

CCC32

CCA148

CCG9

Thr T9.3%

ACU69

ACC129

ACA152

ACG1

Ala A6.6%

GCU51

GCC101

GCA91

GCG6

Tyr Y3.3%

UAU44

UAC82

His H2.6%

CAU42

CAC58

Gln Q2.6%

CAA97

CAG3

Asn N4.0%

AAU46

AAC104

Lys K2.4%

AAA84

AAG6

Asp D1.7%

GAU28

GAC37

Glu E2.4%

GAA77

GAG13

Cys C0.8%

UGU18

UGC14

Trp W2.9%

UGA96

UGG15

Arg R1.8%

CGU5

CGC14

CGA49

CGG1

Gly G5.8%

GGU27

GGC68

GGA99

GGG27

Stop 0.3%

UAA6

UAG2

AGA0

AGG2

Genome map

38 annotated features in NC_007698.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA22
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (38.3%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 38 features as a table

Annotated features of Testudo marginata (NC_007698.1)
Feature	Type	Start	End	Length	Strand	GC
tRNA-Phe	Transfer RNA	1	69	69 bp	+	–
12S rRNA	Ribosomal RNA	70	1,037	968 bp	+	41.3%
tRNA-Val	Transfer RNA	1,038	1,108	71 bp	+	–
16S rRNA	Ribosomal RNA	1,109	2,705	1,597 bp	+	38.7%
tRNA-Leu (UUR)	Transfer RNA	2,706	2,781	76 bp	+	–
ND1	Protein-coding gene	2,782	3,753	972 bp	+	40.4%
tRNA-Ile	Transfer RNA	3,753	3,822	70 bp	+	–
tRNA-Gln	Transfer RNA	3,822	3,892	71 bp	−	–
tRNA-Met	Transfer RNA	3,892	3,960	69 bp	+	–
ND2	Protein-coding gene	3,965	5,005	1,041 bp	+	39.4%
tRNA-Trp	Transfer RNA	5,004	5,081	78 bp	+	–
tRNA-Ala	Transfer RNA	5,083	5,151	69 bp	−	–
tRNA-Asn	Transfer RNA	5,154	5,226	73 bp	−	–
tRNA-Cys	Transfer RNA	5,254	5,319	66 bp	−	–
tRNA-Tyr	Transfer RNA	5,320	5,390	71 bp	−	–
COX1	Protein-coding gene	5,392	6,939	1,548 bp	+	41.3%
tRNA-Ser (UCN)	Transfer RNA	6,931	7,001	71 bp	+	–
tRNA-Asp	Transfer RNA	7,002	7,071	70 bp	+	–
COX2	Protein-coding gene	7,072	7,758	687 bp	+	38.9%
tRNA-Lys	Transfer RNA	7,765	7,835	71 bp	+	–
ATP8	Protein-coding gene	7,837	8,010	174 bp	+	38.5%
ATP6	Protein-coding gene	8,001	8,684	684 bp	+	36.3%
COX3	Protein-coding gene	8,684	9,467	784 bp	+	41.7%
tRNA-Gly	Transfer RNA	9,468	9,536	69 bp	+	–
ND3	Protein-coding gene	9,537	9,888	352 bp	+	39.3%
tRNA-Arg	Transfer RNA	9,888	9,956	69 bp	+	–
ND4L	Protein-coding gene	9,957	10,253	297 bp	+	40.7%
ND4	Protein-coding gene	10,247	11,624	1,378 bp	+	38.5%
tRNA-His	Transfer RNA	11,625	11,692	68 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	11,693	11,758	66 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	11,758	11,828	71 bp	+	–
ND5	Protein-coding gene	11,831	13,621	1,791 bp	+	39.7%
ND6	Protein-coding gene	13,617	14,141	525 bp	−	39.1%
tRNA-Glu	Transfer RNA	14,142	14,208	67 bp	−	–
CYTB	Protein-coding gene	14,213	15,356	1,144 bp	+	41.6%
tRNA-Thr	Transfer RNA	15,357	15,426	70 bp	+	–
tRNA-Pro	Transfer RNA	15,428	15,495	68 bp	+	–
D-loop	Control region	15,496	19,438	3,943 bp	–	32.3%