Thymallus nigrescens
Kosogol grayling
GC content
44.56%
Whole genome
Non-coding
1,071 bp
D-loop 1,003 bp
Where Kosogol grayling sits among 2,335 species
Whole-genome GC content against maximum lifespan. Kosogol grayling has the 1,113th longest lifespan of them.
- This species
- Other ray-finned fishes
- Other classes
Base composition
Whole genome16,661 bp
- GC skew (G − C) / (G + C)
- −0.234
- AT skew (A − T) / (A + T)
- +0.016
About this record
- Lifespan source
- FishBase v26.06
- Based on
- an observed animal, origin not stated
- Data quality
- acceptable
- Licence
- CC BY-NC 4.0
- Status
- New in build 2
- In the database since
- build 2 (2026-11-13)
The lifespan value is for non-commercial use only; the genome data is not restricted by it.
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome16,661 bp
Protein-coding11,409 bp
12S rRNA947 bp
16S rRNA1,679 bp
All tRNA1,555 bp
D-loop1,003 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 975 bp | 46.26% | −0.282 | −0.095 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,048 bp | 43.03% | −0.388 | −0.008 | |||||
| COX1Cytochrome c oxidase subunit I | 1,551 bp | 44.62% | −0.176 | −0.129 | |||||
| COX2Cytochrome c oxidase subunit II | 691 bp | 43.71% | −0.252 | −0.044 | |||||
| ATP8ATP synthase F0 subunit 8 | 168 bp | 48.21% | −0.383 | +0.057 | |||||
| ATP6ATP synthase F0 subunit 6 | 683 bp | 42.90% | −0.386 | −0.077 | |||||
| COX3Cytochrome c oxidase subunit III | 785 bp | 45.48% | −0.261 | −0.089 | |||||
| ND3NADH dehydrogenase subunit 3 | 349 bp | 44.99% | −0.312 | −0.135 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 48.15% | −0.245 | −0.078 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,381 bp | 42.94% | −0.282 | −0.036 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,839 bp | 42.80% | −0.339 | −0.004 | |||||
| ND6NADH dehydrogenase subunit 6 | 522 bp | 48.28% | −0.444 | +0.474 | |||||
| CYTBCytochrome b | 1,141 bp | 44.26% | −0.307 | −0.050 | |||||
| 12S rRNA12S ribosomal RNA | 947 bp | 50.05% | −0.084 | +0.192 | |||||
| 16S rRNA16S ribosomal RNA | 1,679 bp | 47.59% | −0.049 | +0.241 | |||||
| D-loopControl region (D-loop) | 1,003 bp | 34.40% | −0.177 | −0.009 |
Codon usage
Counts per codon, darker is more frequent3,807 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.0%
UUU134
UUC94
Leu L17.4%
UUA116
UUG28
CUU199
CUC99
CUA173
CUG46
Ile I6.8%
AUU157
AUC102
Met M4.0%
AUA93
AUG60
Val V6.2%
GUU68
GUC49
GUA86
GUG33
Ser S6.3%
UCU57
UCC59
UCA59
UCG7
AGU11
AGC46
Pro P5.6%
CCU55
CCC78
CCA74
CCG7
Thr T8.1%
ACU64
ACC120
ACA114
ACG11
Ala A9.1%
GCU84
GCC143
GCA106
GCG15
Tyr Y2.9%
UAU56
UAC54
His H2.7%
CAU23
CAC81
Gln Q2.6%
CAA78
CAG21
Asn N3.0%
AAU51
AAC62
Lys K2.0%
AAA66
AAG10
Asp D2.0%
GAU29
GAC48
Glu E2.8%
GAA76
GAG29
Cys C0.7%
UGU10
UGC17
Trp W3.2%
UGA103
UGG19
Arg R2.0%
CGU10
CGC10
CGA47
CGG9
Gly G6.4%
GGU50
GGC56
GGA90
GGG49
Stop 0.2%
UAA5
UAG1
AGA0
AGG0
Genome map
38 annotated features in NC_056307.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (44.6%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 68 | 68 bp | + | – |
| 12S rRNA | Ribosomal RNA | 69 | 1,015 | 947 bp | + | 50.1% |
| tRNA-Val | Transfer RNA | 1,016 | 1,087 | 72 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,088 | 2,766 | 1,679 bp | + | 47.6% |
| tRNA-Leu (UUR) | Transfer RNA | 2,767 | 2,841 | 75 bp | + | – |
| ND1 | Protein-coding gene | 2,842 | 3,816 | 975 bp | + | 46.3% |
| tRNA-Ile | Transfer RNA | 3,825 | 3,896 | 72 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,894 | 3,964 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 3,964 | 4,032 | 69 bp | + | – |
| ND2 | Protein-coding gene | 4,033 | 5,080 | 1,048 bp | + | 43.0% |
| tRNA-Trp | Transfer RNA | 5,081 | 5,152 | 72 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,154 | 5,222 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,224 | 5,296 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,330 | 5,396 | 67 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,397 | 5,467 | 71 bp | − | – |
| COX1 | Protein-coding gene | 5,469 | 7,019 | 1,551 bp | + | 44.6% |
| tRNA-Ser (UCN) | Transfer RNA | 7,020 | 7,090 | 71 bp | − | – |
| tRNA-Asp | Transfer RNA | 7,095 | 7,168 | 74 bp | + | – |
| COX2 | Protein-coding gene | 7,183 | 7,873 | 691 bp | + | 43.7% |
| tRNA-Lys | Transfer RNA | 7,874 | 7,947 | 74 bp | + | – |
| ATP8 | Protein-coding gene | 7,949 | 8,116 | 168 bp | + | 48.2% |
| ATP6 | Protein-coding gene | 8,107 | 8,789 | 683 bp | + | 42.9% |
| COX3 | Protein-coding gene | 8,790 | 9,574 | 785 bp | + | 45.5% |
| tRNA-Gly | Transfer RNA | 9,575 | 9,644 | 70 bp | + | – |
| ND3 | Protein-coding gene | 9,645 | 9,993 | 349 bp | + | 45.0% |
| tRNA-Arg | Transfer RNA | 9,994 | 10,063 | 70 bp | + | – |
| ND4L | Protein-coding gene | 10,064 | 10,360 | 297 bp | + | 48.2% |
| ND4 | Protein-coding gene | 10,354 | 11,734 | 1,381 bp | + | 42.9% |
| tRNA-His | Transfer RNA | 11,735 | 11,803 | 69 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,804 | 11,872 | 69 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,874 | 11,946 | 73 bp | + | – |
| ND5 | Protein-coding gene | 11,947 | 13,785 | 1,839 bp | + | 42.8% |
| ND6 | Protein-coding gene | 13,782 | 14,303 | 522 bp | − | 48.3% |
| tRNA-Glu | Transfer RNA | 14,304 | 14,372 | 69 bp | − | – |
| CYTB | Protein-coding gene | 14,377 | 15,517 | 1,141 bp | + | 44.3% |
| tRNA-Thr | Transfer RNA | 15,518 | 15,589 | 72 bp | + | – |
| tRNA-Pro | Transfer RNA | 15,589 | 15,658 | 70 bp | − | – |
| D-loop | Control region | 15,659 | 16,661 | 1,003 bp | – | 34.4% |