Thymallus thymallus
Grayling
GC content
44.85%
Whole genome
Non-coding
1,069 bp
D-loop 1,001 bp
Where Grayling sits among 2,335 species
Whole-genome GC content against maximum lifespan. Grayling has the 1,292nd longest lifespan of them.
- This species
- Other ray-finned fishes
- Other classes
Base composition
Whole genome16,657 bp
- GC skew (G − C) / (G + C)
- −0.228
- AT skew (A − T) / (A + T)
- +0.018
About this record
- Lifespan source
- FishBase v26.06
- Based on
- an animal in the wild
- Data quality
- acceptable
- Licence
- CC BY-NC 4.0
- Status
- New in build 2
- In the database since
- build 2 (2026-11-13)
The lifespan value is for non-commercial use only; the genome data is not restricted by it.
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome16,657 bp
Protein-coding11,411 bp
12S rRNA946 bp
16S rRNA1,678 bp
All tRNA1,553 bp
D-loop1,001 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 975 bp | 45.44% | −0.287 | −0.068 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,050 bp | 44.10% | −0.378 | −0.019 | |||||
| COX1Cytochrome c oxidase subunit I | 1,551 bp | 45.20% | −0.161 | −0.134 | |||||
| COX2Cytochrome c oxidase subunit II | 691 bp | 44.14% | −0.252 | −0.031 | |||||
| ATP8ATP synthase F0 subunit 8 | 168 bp | 47.62% | −0.400 | +0.068 | |||||
| ATP6ATP synthase F0 subunit 6 | 684 bp | 41.96% | −0.394 | −0.068 | |||||
| COX3Cytochrome c oxidase subunit III | 785 bp | 45.99% | −0.258 | −0.090 | |||||
| ND3NADH dehydrogenase subunit 3 | 349 bp | 42.98% | −0.280 | −0.126 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 49.83% | −0.243 | −0.101 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,381 bp | 43.30% | −0.278 | −0.034 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,839 bp | 43.07% | −0.328 | +0.001 | |||||
| ND6NADH dehydrogenase subunit 6 | 522 bp | 47.89% | −0.432 | +0.478 | |||||
| CYTBCytochrome b | 1,141 bp | 45.14% | −0.282 | −0.061 | |||||
| 12S rRNA12S ribosomal RNA | 946 bp | 50.32% | −0.080 | +0.191 | |||||
| 16S rRNA16S ribosomal RNA | 1,678 bp | 47.91% | −0.052 | +0.236 | |||||
| D-loopControl region (D-loop) | 1,001 bp | 35.56% | −0.157 | +0.005 |
Codon usage
Counts per codon, darker is more frequent3,809 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.0%
UUU127
UUC101
Leu L17.3%
UUA107
UUG34
CUU191
CUC107
CUA176
CUG43
Ile I6.8%
AUU163
AUC96
Met M4.0%
AUA94
AUG59
Val V6.2%
GUU73
GUC41
GUA92
GUG31
Ser S6.2%
UCU57
UCC53
UCA64
UCG9
AGU13
AGC42
Pro P5.6%
CCU54
CCC77
CCA74
CCG8
Thr T7.9%
ACU62
ACC114
ACA109
ACG16
Ala A9.3%
GCU84
GCC147
GCA112
GCG13
Tyr Y2.9%
UAU48
UAC62
His H2.7%
CAU22
CAC82
Gln Q2.6%
CAA79
CAG19
Asn N3.0%
AAU41
AAC73
Lys K2.0%
AAA64
AAG12
Asp D2.0%
GAU33
GAC45
Glu E2.8%
GAA79
GAG27
Cys C0.7%
UGU11
UGC16
Trp W3.2%
UGA95
UGG26
Arg R2.1%
CGU10
CGC11
CGA48
CGG10
Gly G6.4%
GGU56
GGC51
GGA84
GGG54
Stop 0.2%
UAA7
UAG1
AGA0
AGG0
Genome map
38 annotated features in NC_012928.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (44.8%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| D-loop | Control region | 1 | 1,001 | 1,001 bp | – | 35.6% |
| tRNA-Phe | Transfer RNA | 1,002 | 1,069 | 68 bp | + | – |
| 12S rRNA | Ribosomal RNA | 1,070 | 2,015 | 946 bp | + | 50.3% |
| tRNA-Val | Transfer RNA | 2,016 | 2,087 | 72 bp | + | – |
| 16S rRNA | Ribosomal RNA | 2,088 | 3,765 | 1,678 bp | + | 47.9% |
| tRNA-Leu (UUR) | Transfer RNA | 3,766 | 3,840 | 75 bp | + | – |
| ND1 | Protein-coding gene | 3,841 | 4,815 | 975 bp | + | 45.4% |
| tRNA-Ile | Transfer RNA | 4,824 | 4,895 | 72 bp | + | – |
| tRNA-Gln | Transfer RNA | 4,893 | 4,963 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 4,963 | 5,031 | 69 bp | + | – |
| ND2 | Protein-coding gene | 5,032 | 6,081 | 1,050 bp | + | 44.1% |
| tRNA-Trp | Transfer RNA | 6,082 | 6,151 | 70 bp | + | – |
| tRNA-Ala | Transfer RNA | 6,154 | 6,222 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 6,224 | 6,296 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 6,330 | 6,396 | 67 bp | − | – |
| tRNA-Tyr | Transfer RNA | 6,397 | 6,467 | 71 bp | − | – |
| COX1 | Protein-coding gene | 6,469 | 8,019 | 1,551 bp | + | 45.2% |
| tRNA-Ser (UCN) | Transfer RNA | 8,020 | 8,090 | 71 bp | − | – |
| tRNA-Asp | Transfer RNA | 8,094 | 8,166 | 73 bp | + | – |
| COX2 | Protein-coding gene | 8,181 | 8,871 | 691 bp | + | 44.1% |
| tRNA-Lys | Transfer RNA | 8,872 | 8,945 | 74 bp | + | – |
| ATP8 | Protein-coding gene | 8,947 | 9,114 | 168 bp | + | 47.6% |
| ATP6 | Protein-coding gene | 9,105 | 9,788 | 684 bp | + | 42.0% |
| COX3 | Protein-coding gene | 9,788 | 10,572 | 785 bp | + | 46.0% |
| tRNA-Gly | Transfer RNA | 10,573 | 10,642 | 70 bp | + | – |
| ND3 | Protein-coding gene | 10,643 | 10,991 | 349 bp | + | 43.0% |
| tRNA-Arg | Transfer RNA | 10,992 | 11,061 | 70 bp | + | – |
| ND4L | Protein-coding gene | 11,062 | 11,358 | 297 bp | + | 49.8% |
| ND4 | Protein-coding gene | 11,352 | 12,732 | 1,381 bp | + | 43.3% |
| tRNA-His | Transfer RNA | 12,733 | 12,801 | 69 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 12,802 | 12,871 | 70 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 12,873 | 12,945 | 73 bp | + | – |
| ND5 | Protein-coding gene | 12,946 | 14,784 | 1,839 bp | + | 43.1% |
| ND6 | Protein-coding gene | 14,781 | 15,302 | 522 bp | − | 47.9% |
| tRNA-Glu | Transfer RNA | 15,303 | 15,371 | 69 bp | − | – |
| CYTB | Protein-coding gene | 15,376 | 16,516 | 1,141 bp | + | 45.1% |
| tRNA-Thr | Transfer RNA | 16,517 | 16,588 | 72 bp | + | – |
| tRNA-Pro | Transfer RNA | 16,588 | 16,657 | 70 bp | − | – |