Thymallus tugarinae
lower Amur grayling
GC content
45.14%
Whole genome
Non-coding
1,070 bp
D-loop 1,002 bp
Where lower Amur grayling sits among 2,335 species
Whole-genome GC content against maximum lifespan. lower Amur grayling has the 1,945th longest lifespan of them.
- This species
- Other ray-finned fishes
- Other classes
Base composition
Whole genome16,662 bp
- GC skew (G − C) / (G + C)
- −0.225
- AT skew (A − T) / (A + T)
- +0.016
About this record
- Lifespan source
- FishBase v26.06
- Based on
- an animal in the wild
- Data quality
- acceptable
- Licence
- CC BY-NC 4.0
- Status
- New in build 2
- In the database since
- build 2 (2026-11-13)
The lifespan value is for non-commercial use only; the genome data is not restricted by it.
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome16,662 bp
Protein-coding11,410 bp
12S rRNA947 bp
16S rRNA1,679 bp
All tRNA1,556 bp
D-loop1,002 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 975 bp | 45.95% | −0.268 | −0.089 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,049 bp | 44.52% | −0.349 | +0.003 | |||||
| COX1Cytochrome c oxidase subunit I | 1,551 bp | 45.33% | −0.175 | −0.127 | |||||
| COX2Cytochrome c oxidase subunit II | 691 bp | 45.15% | −0.250 | −0.040 | |||||
| ATP8ATP synthase F0 subunit 8 | 168 bp | 47.02% | −0.418 | +0.079 | |||||
| ATP6ATP synthase F0 subunit 6 | 684 bp | 42.98% | −0.374 | −0.067 | |||||
| COX3Cytochrome c oxidase subunit III | 785 bp | 46.12% | −0.265 | −0.078 | |||||
| ND3NADH dehydrogenase subunit 3 | 349 bp | 44.41% | −0.290 | −0.134 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 48.49% | −0.250 | −0.111 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,381 bp | 44.24% | −0.283 | −0.031 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,839 bp | 43.01% | −0.315 | −0.019 | |||||
| ND6NADH dehydrogenase subunit 6 | 522 bp | 48.08% | −0.442 | +0.476 | |||||
| CYTBCytochrome b | 1,141 bp | 45.75% | −0.280 | −0.060 | |||||
| 12S rRNA12S ribosomal RNA | 947 bp | 50.26% | −0.084 | +0.185 | |||||
| 16S rRNA16S ribosomal RNA | 1,679 bp | 47.71% | −0.046 | +0.237 | |||||
| D-loopControl region (D-loop) | 1,002 bp | 35.53% | −0.163 | −0.009 |
Codon usage
Counts per codon, darker is more frequent3,808 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.0%
UUU127
UUC100
Leu L17.2%
UUA107
UUG29
CUU184
CUC101
CUA172
CUG63
Ile I6.9%
AUU161
AUC101
Met M4.0%
AUA83
AUG69
Val V6.3%
GUU69
GUC48
GUA85
GUG36
Ser S6.2%
UCU59
UCC57
UCA61
UCG6
AGU11
AGC43
Pro P5.6%
CCU46
CCC81
CCA76
CCG10
Thr T8.0%
ACU65
ACC116
ACA111
ACG11
Ala A9.3%
GCU88
GCC143
GCA113
GCG12
Tyr Y2.9%
UAU52
UAC58
His H2.8%
CAU21
CAC84
Gln Q2.6%
CAA82
CAG16
Asn N3.0%
AAU43
AAC72
Lys K2.0%
AAA59
AAG17
Asp D2.0%
GAU28
GAC49
Glu E2.8%
GAA76
GAG29
Cys C0.7%
UGU11
UGC16
Trp W3.2%
UGA97
UGG26
Arg R2.0%
CGU10
CGC8
CGA48
CGG11
Gly G6.4%
GGU54
GGC55
GGA78
GGG57
Stop 0.2%
UAA5
UAG2
AGA0
AGG0
Genome map
38 annotated features in NC_027409.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- D-loop1
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (45.1%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1 | 68 | 68 bp | + | – |
| 12S rRNA | Ribosomal RNA | 69 | 1,015 | 947 bp | + | 50.3% |
| tRNA-Val | Transfer RNA | 1,016 | 1,087 | 72 bp | + | – |
| 16S rRNA | Ribosomal RNA | 1,088 | 2,766 | 1,679 bp | + | 47.7% |
| tRNA-Leu (UUR) | Transfer RNA | 2,767 | 2,841 | 75 bp | + | – |
| ND1 | Protein-coding gene | 2,842 | 3,816 | 975 bp | + | 46.0% |
| tRNA-Ile | Transfer RNA | 3,825 | 3,896 | 72 bp | + | – |
| tRNA-Gln | Transfer RNA | 3,894 | 3,964 | 71 bp | − | – |
| tRNA-Met | Transfer RNA | 3,964 | 4,032 | 69 bp | + | – |
| ND2 | Protein-coding gene | 4,033 | 5,081 | 1,049 bp | + | 44.5% |
| tRNA-Trp | Transfer RNA | 5,082 | 5,153 | 72 bp | + | – |
| tRNA-Ala | Transfer RNA | 5,155 | 5,223 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 5,225 | 5,297 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 5,331 | 5,397 | 67 bp | − | – |
| tRNA-Tyr | Transfer RNA | 5,398 | 5,468 | 71 bp | − | – |
| COX1 | Protein-coding gene | 5,470 | 7,020 | 1,551 bp | + | 45.3% |
| tRNA-Ser (UCN) | Transfer RNA | 7,021 | 7,091 | 71 bp | − | – |
| tRNA-Asp | Transfer RNA | 7,096 | 7,169 | 74 bp | + | – |
| COX2 | Protein-coding gene | 7,184 | 7,874 | 691 bp | + | 45.2% |
| tRNA-Lys | Transfer RNA | 7,875 | 7,948 | 74 bp | + | – |
| ATP8 | Protein-coding gene | 7,950 | 8,117 | 168 bp | + | 47.0% |
| ATP6 | Protein-coding gene | 8,108 | 8,791 | 684 bp | + | 43.0% |
| COX3 | Protein-coding gene | 8,791 | 9,575 | 785 bp | + | 46.1% |
| tRNA-Gly | Transfer RNA | 9,576 | 9,645 | 70 bp | + | – |
| ND3 | Protein-coding gene | 9,646 | 9,994 | 349 bp | + | 44.4% |
| tRNA-Arg | Transfer RNA | 9,995 | 10,064 | 70 bp | + | – |
| ND4L | Protein-coding gene | 10,065 | 10,361 | 297 bp | + | 48.5% |
| ND4 | Protein-coding gene | 10,355 | 11,735 | 1,381 bp | + | 44.2% |
| tRNA-His | Transfer RNA | 11,736 | 11,804 | 69 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 11,805 | 11,874 | 70 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 11,876 | 11,948 | 73 bp | + | – |
| ND5 | Protein-coding gene | 11,949 | 13,787 | 1,839 bp | + | 43.0% |
| ND6 | Protein-coding gene | 13,784 | 14,305 | 522 bp | − | 48.1% |
| tRNA-Glu | Transfer RNA | 14,306 | 14,374 | 69 bp | − | – |
| CYTB | Protein-coding gene | 14,379 | 15,519 | 1,141 bp | + | 45.8% |
| tRNA-Thr | Transfer RNA | 15,520 | 15,591 | 72 bp | + | – |
| tRNA-Pro | Transfer RNA | 15,591 | 15,660 | 70 bp | − | – |
| D-loop | Control region | 15,661 | 16,662 | 1,002 bp | – | 35.5% |