Ursus arctos
Brown bear
GC content
41.31%
Whole genome
Non-coding
1,638 bp
Including the D-loop
Where Brown bear sits among 2,335 species
Whole-genome GC content against maximum lifespan. Brown bear has the 254th longest lifespan of them.
- This species
- Other mammals
- Other classes
Base composition
Whole genome17,020 bp
- GC skew (G − C) / (G + C)
- −0.239
- AT skew (A − T) / (A + T)
- +0.053
Life history
- Maximum lifespan
- 40 years
- Adult weight
- 277.5 kg
- Birth weight
- 510 g
- Body temperature
- 37.3 °C
- Female sexual maturity
- 3.6 years
- Male sexual maturity
- 3.6 years
- Gestation or incubation
- 103 days
- Litter or clutch size
- 2
- Litters per year
- 0.4
Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 02252)
About this record
- Lifespan source
- AnAge build 15
- Based on
- an animal in captivity
- Data quality
- acceptable, large sample
- Licence
- Free to use, subject to the Human Ageing Genomic Resources conditions of use
- Status
- Updated in build 2
- In the database since
- build 1 (2015-07-01)
- Last changed
- build 2 (2026-11-13)
Every change against the 2015 build is in the changelog download.
Composition by region
Reference (L) strand as deposited in RefSeqWhole genome17,020 bp
Protein-coding11,341 bp
12S rRNA962 bp
16S rRNA1,579 bp
All tRNA1,508 bp
Genes and regions
Select a column to sort; select a gene to compare it across species| Composition | |||||||||
|---|---|---|---|---|---|---|---|---|---|
| ND1NADH dehydrogenase subunit 1 | 957 bp | 41.38% | −0.359 | +0.005 | |||||
| ND2NADH dehydrogenase subunit 2 | 1,044 bp | 38.89% | −0.448 | +0.107 | |||||
| COX1Cytochrome c oxidase subunit I | 1,545 bp | 43.04% | −0.113 | −0.091 | |||||
| COX2Cytochrome c oxidase subunit II | 684 bp | 40.94% | −0.150 | +0.045 | |||||
| ATP8ATP synthase F0 subunit 8 | 204 bp | 36.77% | −0.413 | +0.147 | |||||
| ATP6ATP synthase F0 subunit 6 | 681 bp | 41.41% | −0.277 | −0.073 | |||||
| COX3Cytochrome c oxidase subunit III | 784 bp | 42.86% | −0.250 | −0.054 | |||||
| ND3NADH dehydrogenase subunit 3 | 346 bp | 42.49% | −0.293 | −0.035 | |||||
| ND4LNADH dehydrogenase subunit 4L | 297 bp | 41.08% | −0.246 | −0.120 | |||||
| ND4NADH dehydrogenase subunit 4 | 1,378 bp | 41.07% | −0.353 | +0.012 | |||||
| ND5NADH dehydrogenase subunit 5 | 1,821 bp | 40.03% | −0.383 | +0.059 | |||||
| ND6NADH dehydrogenase subunit 6 | 528 bp | 35.99% | −0.495 | +0.314 | |||||
| CYTBCytochrome b | 1,140 bp | 44.30% | −0.347 | +0.033 | |||||
| 12S rRNA12S ribosomal RNA | 962 bp | 42.41% | −0.083 | +0.199 | |||||
| 16S rRNA16S ribosomal RNA | 1,579 bp | 39.39% | −0.074 | +0.197 |
Codon usage
Counts per codon, darker is more frequent3,802 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).
Phe F6.3%
UUU123
UUC115
Leu L15.9%
UUA87
UUG27
CUU71
CUC93
CUA255
CUG70
Ile I8.4%
AUU163
AUC157
Met M6.0%
AUA160
AUG70
Val V5.4%
GUU39
GUC49
GUA92
GUG27
Ser S7.4%
UCU53
UCC70
UCA86
UCG11
AGU27
AGC35
Pro P5.2%
CCU67
CCC59
CCA59
CCG12
Thr T8.0%
ACU87
ACC75
ACA128
ACG16
Ala A7.1%
GCU66
GCC90
GCA98
GCG15
Tyr Y3.5%
UAU67
UAC67
His H2.6%
CAU42
CAC55
Gln Q2.3%
CAA71
CAG17
Asn N3.7%
AAU66
AAC74
Lys K2.6%
AAA80
AAG20
Asp D1.8%
GAU30
GAC40
Glu E2.7%
GAA68
GAG33
Cys C0.7%
UGU9
UGC16
Trp W2.7%
UGA89
UGG15
Arg R1.7%
CGU4
CGC11
CGA41
CGG9
Gly G5.7%
GGU28
GGC49
GGA104
GGG35
Stop 0.3%
UAA6
UAG3
AGA1
AGG0
Genome map
37 annotated features in NC_003427.1Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.
- Protein-coding13
- rRNA2
- tRNA22
- Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
- Middle ring: GC% of each gene against the whole genome (41.3%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 37 features as a table
| Feature | Type | Start | End | Length | Strand | GC |
|---|---|---|---|---|---|---|
| tRNA-Phe | Transfer RNA | 1,139 | 1,206 | 68 bp | + | – |
| 12S rRNA | Ribosomal RNA | 1,207 | 2,168 | 962 bp | + | 42.4% |
| tRNA-Val | Transfer RNA | 2,169 | 2,234 | 66 bp | + | – |
| 16S rRNA | Ribosomal RNA | 2,235 | 3,813 | 1,579 bp | + | 39.4% |
| tRNA-Leu (UUR) | Transfer RNA | 3,815 | 3,889 | 75 bp | + | – |
| ND1 | Protein-coding gene | 3,892 | 4,848 | 957 bp | + | 41.4% |
| tRNA-Ile | Transfer RNA | 4,847 | 4,916 | 70 bp | + | – |
| tRNA-Gln | Transfer RNA | 4,914 | 4,986 | 73 bp | − | – |
| tRNA-Met | Transfer RNA | 4,988 | 5,056 | 69 bp | + | – |
| ND2 | Protein-coding gene | 5,057 | 6,100 | 1,044 bp | + | 38.9% |
| tRNA-Trp | Transfer RNA | 6,099 | 6,166 | 68 bp | + | – |
| tRNA-Ala | Transfer RNA | 6,175 | 6,243 | 69 bp | − | – |
| tRNA-Asn | Transfer RNA | 6,244 | 6,316 | 73 bp | − | – |
| tRNA-Cys | Transfer RNA | 6,350 | 6,416 | 67 bp | − | – |
| tRNA-Tyr | Transfer RNA | 6,416 | 6,482 | 67 bp | − | – |
| COX1 | Protein-coding gene | 6,484 | 8,028 | 1,545 bp | + | 43.0% |
| tRNA-Ser (UCN) | Transfer RNA | 8,025 | 8,096 | 72 bp | − | – |
| tRNA-Asp | Transfer RNA | 8,101 | 8,167 | 67 bp | + | – |
| COX2 | Protein-coding gene | 8,168 | 8,851 | 684 bp | + | 40.9% |
| tRNA-Lys | Transfer RNA | 8,855 | 8,922 | 68 bp | + | – |
| ATP8 | Protein-coding gene | 8,925 | 9,128 | 204 bp | + | 36.8% |
| ATP6 | Protein-coding gene | 9,086 | 9,766 | 681 bp | + | 41.4% |
| COX3 | Protein-coding gene | 9,766 | 10,549 | 784 bp | + | 42.9% |
| tRNA-Gly | Transfer RNA | 10,550 | 10,618 | 69 bp | + | – |
| ND3 | Protein-coding gene | 10,619 | 10,964 | 346 bp | + | 42.5% |
| tRNA-Arg | Transfer RNA | 10,966 | 11,034 | 69 bp | + | – |
| ND4L | Protein-coding gene | 11,035 | 11,331 | 297 bp | + | 41.1% |
| ND4 | Protein-coding gene | 11,325 | 12,702 | 1,378 bp | + | 41.1% |
| tRNA-His | Transfer RNA | 12,703 | 12,771 | 69 bp | + | – |
| tRNA-Ser (AGY) | Transfer RNA | 12,772 | 12,830 | 59 bp | + | – |
| tRNA-Leu (CUN) | Transfer RNA | 12,831 | 12,900 | 70 bp | + | – |
| ND5 | Protein-coding gene | 12,901 | 14,721 | 1,821 bp | + | 40.0% |
| ND6 | Protein-coding gene | 14,705 | 15,232 | 528 bp | − | 36.0% |
| tRNA-Glu | Transfer RNA | 15,233 | 15,301 | 69 bp | − | – |
| CYTB | Protein-coding gene | 15,306 | 16,445 | 1,140 bp | + | 44.3% |
| tRNA-Thr | Transfer RNA | 16,446 | 16,515 | 70 bp | + | – |
| tRNA-Pro | Transfer RNA | 16,516 | 16,580 | 65 bp | − | – |