MitoAge

Varecia variegata

Black-and-white ruffed lemur, or Ruffed lemur

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Maximum lifespan
40 yr
AnAge record
mtDNA size
16,965 bp
RefSeq NC_012773.1
GC content
38.06%
Whole genome
Non-coding
1,577 bp
D-loop 1,490 bp

Where Black-and-white ruffed lemur, or Ruffed lemur sits among 2,335 species

Whole-genome GC content against maximum lifespan. Black-and-white ruffed lemur, or Ruffed lemur has the 258th longest lifespan of them.

10203040501 yr10 yr100 yrMaximum lifespan, log scaleGC %Black-and-white ruffed lemur, or Ruffed lemur
  • This species
  • Other mammals
  • Other classes
Plot other mtDNA features with this species highlighted

Base composition

Whole genome16,965 bp
GC skew (G − C) / (G + C)
−0.357
AT skew (A − T) / (A + T)
+0.074

Life history

Maximum lifespan
40 years
Adult weight
3.5 kg
Birth weight
107.2 g
Female sexual maturity
605 days
Male sexual maturity
649 days
Gestation or incubation
98 days
Litter or clutch size
1.9
Litters per year
1

Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 02816)

About this record

Lifespan source
AnAge build 15
Based on
an animal in captivity
Data quality
high, large sample
Licence
Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status
Updated in build 2
In the database since
build 1 (2015-07-01)
Last changed
build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq
Whole genome16,965 bp
Protein-coding11,339 bp
12S rRNA957 bp
16S rRNA1,579 bp
All tRNA1,517 bp
D-loop1,490 bp

Genes and regions

Select a column to sort; select a gene to compare it across species
Composition
ND1NADH dehydrogenase subunit 1957 bp30.6%29.8%11.1%28.5%39.60%−0.441+0.014
ND2NADH dehydrogenase subunit 21,044 bp35.2%29.6%8.0%27.2%35.25%−0.543+0.086
COX1Cytochrome c oxidase subunit I1,542 bp28.1%31.8%15.7%24.4%40.08%−0.217−0.063
COX2Cytochrome c oxidase subunit II684 bp32.3%28.9%12.9%25.9%38.74%−0.336+0.055
ATP8ATP synthase F0 subunit 8207 bp36.7%30.4%6.8%26.1%32.85%−0.588+0.094
ATP6ATP synthase F0 subunit 6681 bp31.0%32.0%9.7%27.3%37.00%−0.476−0.016
COX3Cytochrome c oxidase subunit III784 bp26.5%31.1%14.0%28.3%42.35%−0.337−0.080
ND3NADH dehydrogenase subunit 3348 bp33.0%29.3%10.1%27.6%37.64%−0.466+0.060
ND4LNADH dehydrogenase subunit 4L297 bp30.0%32.7%10.1%27.3%37.37%−0.459−0.043
ND4NADH dehydrogenase subunit 41,378 bp33.0%30.7%9.8%26.5%36.28%−0.460+0.036
ND5NADH dehydrogenase subunit 51,812 bp34.1%29.5%9.5%26.9%36.42%−0.479+0.073
ND6NADH dehydrogenase subunit 6519 bp44.1%20.0%6.0%29.9%35.84%−0.667+0.375
CYTBCytochrome b1,140 bp29.3%28.9%12.2%29.6%41.75%−0.416+0.006
12S rRNA12S ribosomal RNA957 bp37.3%23.2%16.4%23.1%39.50%−0.169+0.233
16S rRNA16S ribosomal RNA1,579 bp38.1%24.4%15.6%21.8%37.49%−0.166+0.220
D-loopControl region (D-loop)1,490 bp32.9%28.6%13.2%25.3%38.52%−0.314+0.070

Codon usage

Counts per codon, darker is more frequent

3,797 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F6.2%
UUU110
UUC127
Leu L16.3%
UUA132
UUG13
CUU104
CUC94
CUA266
CUG10
Ile I9.1%
AUU192
AUC153
Met M6.8%
AUA231
AUG26
Val V4.4%
GUU42
GUC33
GUA81
GUG11
Ser S7.5%
UCU61
UCC75
UCA89
UCG6
AGU20
AGC35
Pro P5.1%
CCU64
CCC61
CCA68
CCG0
Thr T8.1%
ACU68
ACC97
ACA134
ACG7
Ala A6.2%
GCU59
GCC94
GCA80
GCG1
Tyr Y3.5%
UAU71
UAC60
His H2.5%
CAU41
CAC55
Gln Q2.4%
CAA84
CAG6
Asn N4.3%
AAU71
AAC92
Lys K2.6%
AAA90
AAG7
Asp D1.7%
GAU34
GAC29
Glu E2.5%
GAA82
GAG14
Cys C0.7%
UGU11
UGC16
Trp W2.8%
UGA98
UGG7
Arg R1.7%
CGU9
CGC14
CGA39
CGG1
Gly G5.6%
GGU54
GGC55
GGA73
GGG29
Stop 0.3%
UAA6
UAG3
AGA1
AGG1

Genome map

38 annotated features in NC_012773.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

  • Protein-coding13
  • rRNA2
  • tRNA22
  • D-loop1
  • Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
  • Middle ring: GC% of each gene against the whole genome (38.1%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).
All 38 features as a table
FeatureTypeStartEndLengthStrandGC
tRNA-PheTransfer RNA17070 bp+–
12S rRNARibosomal RNA711,027957 bp+39.5%
tRNA-ValTransfer RNA1,0281,09669 bp+–
16S rRNARibosomal RNA1,0982,6761,579 bp+37.5%
tRNA-Leu (UUR)Transfer RNA2,6772,75276 bp+–
ND1Protein-coding gene2,7563,712957 bp+39.6%
tRNA-IleTransfer RNA3,7113,77969 bp+–
tRNA-GlnTransfer RNA3,7773,85175 bp−–
tRNA-MetTransfer RNA3,8553,92369 bp+–
ND2Protein-coding gene3,9244,9671,044 bp+35.3%
tRNA-TrpTransfer RNA4,9665,03267 bp+–
tRNA-AlaTransfer RNA5,0415,10969 bp−–
tRNA-AsnTransfer RNA5,1125,18473 bp−–
tRNA-CysTransfer RNA5,2205,28566 bp−–
tRNA-TyrTransfer RNA5,2865,35166 bp−–
COX1Protein-coding gene5,3586,8991,542 bp+40.1%
tRNA-Ser (UCN)Transfer RNA6,9016,97272 bp−–
tRNA-AspTransfer RNA6,9767,04469 bp+–
COX2Protein-coding gene7,0457,728684 bp+38.7%
tRNA-LysTransfer RNA7,7327,79968 bp+–
ATP8Protein-coding gene7,8008,006207 bp+32.9%
ATP6Protein-coding gene7,9618,641681 bp+37.0%
COX3Protein-coding gene8,6419,424784 bp+42.4%
tRNA-GlyTransfer RNA9,4259,49470 bp+–
ND3Protein-coding gene9,4969,843348 bp+37.6%
tRNA-ArgTransfer RNA9,8469,91368 bp+–
ND4LProtein-coding gene9,91510,211297 bp+37.4%
ND4Protein-coding gene10,20511,5821,378 bp+36.3%
tRNA-HisTransfer RNA11,58311,65270 bp+–
tRNA-Ser (AGY)Transfer RNA11,65311,71260 bp+–
tRNA-Leu (CUN)Transfer RNA11,71311,78270 bp+–
ND5Protein-coding gene11,78313,5941,812 bp+36.4%
ND6Protein-coding gene13,60714,125519 bp−35.8%
tRNA-GluTransfer RNA14,12614,19469 bp−–
CYTBProtein-coding gene14,19915,3381,140 bp+41.8%
tRNA-ThrTransfer RNA15,33915,40769 bp+–
tRNA-ProTransfer RNA15,41015,47566 bp−–
D-loopControl region15,47616,9651,490 bp–38.5%

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