Rhizoprionodon acutus

Milk shark

Maximum lifespan

8 yr

AnAge record

mtDNA size

16,693 bp

RefSeq NC_046016.1

GC content

36.99%

Whole genome

Non-coding

1,120 bp

D-loop 1,064 bp

Where Milk shark sits among 2,335 species

Whole-genome GC content against maximum lifespan. Milk shark has the 1,737th longest lifespan of them.

This species
Other sharks and rays
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome16,693 bp

GC skew (G − C) / (G + C): −0.293
AT skew (A − T) / (A + T): +0.004

Life history

Maximum lifespan: 8 years
Adult weight: 2.8 kg

Lifespan from AnAge · AnAge record · other traits from AnAge build 15 (HAGRID 01767)

About this record

Lifespan source: AnAge build 15
Based on: an animal in the wild
Data quality: acceptable, medium sample
Licence: Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status: New in build 2
In the database since: build 2 (2026-11-13)

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome16,693 bp

Protein-coding11,407 bp

12S rRNA951 bp

16S rRNA1,670 bp

All tRNA1,545 bp

D-loop1,064 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Rhizoprionodon acutus genes

ND1NADH dehydrogenase subunit 1	975 bp	29.9%	32.6%	10.8%	26.7%	37.44%	−0.425	−0.043
ND2NADH dehydrogenase subunit 2	1,045 bp	30.8%	31.1%	9.9%	28.2%	38.09%	−0.482	−0.005
COX1Cytochrome c oxidase subunit I	1,557 bp	27.0%	35.8%	15.9%	21.3%	37.19%	−0.143	−0.141
COX2Cytochrome c oxidase subunit II	691 bp	31.8%	31.1%	14.5%	22.6%	37.05%	−0.219	+0.011
ATP8ATP synthase F0 subunit 8	168 bp	38.1%	33.3%	6.0%	22.6%	28.57%	−0.583	+0.067
ATP6ATP synthase F0 subunit 6	684 bp	31.7%	33.6%	9.8%	24.9%	34.65%	−0.435	−0.029
COX3Cytochrome c oxidase subunit III	786 bp	28.6%	31.6%	15.9%	23.9%	39.82%	−0.201	−0.049
ND3NADH dehydrogenase subunit 3	349 bp	24.9%	33.2%	14.9%	26.9%	41.83%	−0.288	−0.143
ND4LNADH dehydrogenase subunit 4L	297 bp	26.3%	33.7%	11.8%	28.3%	40.07%	−0.412	−0.124
ND4NADH dehydrogenase subunit 4	1,381 bp	30.7%	33.2%	10.2%	25.9%	36.13%	−0.435	−0.039
ND5NADH dehydrogenase subunit 5	1,830 bp	31.3%	31.5%	9.9%	27.4%	37.27%	−0.469	−0.003
ND6NADH dehydrogenase subunit 6	522 bp	42.1%	20.3%	8.0%	29.5%	37.55%	−0.571	+0.350
CYTBCytochrome b	1,145 bp	28.6%	33.4%	11.8%	26.1%	37.90%	−0.378	−0.077
12S rRNA12S ribosomal RNA	951 bp	34.4%	25.2%	18.5%	21.9%	40.38%	−0.083	+0.153
16S rRNA16S ribosomal RNA	1,670 bp	37.2%	28.6%	15.9%	18.3%	34.19%	−0.068	+0.130
D-loopControl region (D-loop)	1,064 bp	32.5%	35.3%	12.5%	19.6%	32.14%	−0.222	−0.042

Codon usage

Counts per codon, darker is more frequent

Gene

3,808 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F6.6%

UUU167

UUC83

Leu L17.4%

UUA228

UUG13

CUU141

CUC100

CUA167

CUG14

Ile I9.2%

AUU270

AUC80

Met M4.4%

AUA140

AUG28

Val V4.2%

GUU55

GUC20

GUA74

GUG12

Ser S6.9%

UCU47

UCC69

UCA94

UCG1

AGU22

AGC29

Pro P5.5%

CCU53

CCC62

CCA89

CCG4

Thr T7.8%

ACU65

ACC106

ACA122

ACG4

Ala A7.1%

GCU77

GCC99

GCA89

GCG4

Tyr Y3.2%

UAU90

UAC33

His H2.7%

CAU43

CAC61

Gln Q2.5%

CAA93

CAG3

Asn N3.8%

AAU70

AAC74

Lys K2.2%

AAA84

AAG1

Asp D1.9%

GAU42

GAC31

Glu E2.6%

GAA95

GAG5

Cys C0.7%

UGU14

UGC11

Trp W3.2%

UGA114

UGG8

Arg R1.8%

CGU14

CGC15

CGA37

CGG4

Gly G6.0%

GGU57

GGC58

GGA93

GGG22

Stop 0.2%

UAA7

UAG0

AGA0

AGG1

Genome map

38 annotated features in NC_046016.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA22
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (37.0%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 38 features as a table

Annotated features of Rhizoprionodon acutus (NC_046016.1)
Feature	Type	Start	End	Length	Strand	GC
tRNA-Phe	Transfer RNA	1	70	70 bp	+	–
12S rRNA	Ribosomal RNA	71	1,021	951 bp	+	40.4%
tRNA-Val	Transfer RNA	1,022	1,093	72 bp	+	–
16S rRNA	Ribosomal RNA	1,094	2,763	1,670 bp	+	34.2%
tRNA-Leu (UUR)	Transfer RNA	2,764	2,838	75 bp	+	–
ND1	Protein-coding gene	2,839	3,813	975 bp	+	37.4%
tRNA-Ile	Transfer RNA	3,814	3,883	70 bp	+	–
tRNA-Gln	Transfer RNA	3,885	3,956	72 bp	−	–
tRNA-Met	Transfer RNA	3,957	4,025	69 bp	+	–
ND2	Protein-coding gene	4,026	5,070	1,045 bp	+	38.1%
tRNA-Trp	Transfer RNA	5,071	5,138	68 bp	+	–
tRNA-Ala	Transfer RNA	5,140	5,208	69 bp	−	–
tRNA-Asn	Transfer RNA	5,209	5,281	73 bp	−	–
tRNA-Cys	Transfer RNA	5,316	5,382	67 bp	−	–
tRNA-Tyr	Transfer RNA	5,384	5,452	69 bp	−	–
COX1	Protein-coding gene	5,454	7,010	1,557 bp	+	37.2%
tRNA-Ser (UCN)	Transfer RNA	7,011	7,081	71 bp	−	–
tRNA-Asp	Transfer RNA	7,085	7,153	69 bp	+	–
COX2	Protein-coding gene	7,161	7,851	691 bp	+	37.1%
tRNA-Lys	Transfer RNA	7,852	7,925	74 bp	+	–
ATP8	Protein-coding gene	7,927	8,094	168 bp	+	28.6%
ATP6	Protein-coding gene	8,085	8,768	684 bp	+	34.7%
COX3	Protein-coding gene	8,768	9,553	786 bp	+	39.8%
tRNA-Gly	Transfer RNA	9,556	9,625	70 bp	+	–
ND3	Protein-coding gene	9,626	9,974	349 bp	+	41.8%
tRNA-Arg	Transfer RNA	9,975	10,044	70 bp	+	–
ND4L	Protein-coding gene	10,045	10,341	297 bp	+	40.1%
ND4	Protein-coding gene	10,335	11,715	1,381 bp	+	36.1%
tRNA-His	Transfer RNA	11,716	11,784	69 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	11,785	11,851	67 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	11,852	11,923	72 bp	+	–
ND5	Protein-coding gene	11,925	13,754	1,830 bp	+	37.3%
ND6	Protein-coding gene	13,750	14,271	522 bp	−	37.6%
tRNA-Glu	Transfer RNA	14,272	14,341	70 bp	−	–
CYTB	Protein-coding gene	14,344	15,488	1,145 bp	+	37.9%
tRNA-Thr	Transfer RNA	15,489	15,558	70 bp	+	–
tRNA-Pro	Transfer RNA	15,561	15,629	69 bp	−	–
D-loop	Control region	15,630	16,693	1,064 bp	–	32.1%