Caliraja rhina

Longnose skate

Maximum lifespan

26 yr

AnAge record

mtDNA size

16,910 bp

RefSeq NC_021964.1

GC content

41.44%

Whole genome

Non-coding

1,308 bp

D-loop 1,251 bp

Where Longnose skate sits among 2,335 species

Whole-genome GC content against maximum lifespan. Longnose skate has the 590th longest lifespan of them.

This species
Other sharks and rays
Other classes

Plot other mtDNA features with this species highlighted

Base composition

Whole genome16,910 bp

GC skew (G − C) / (G + C): −0.314
AT skew (A − T) / (A + T): +0.028

Life history

Maximum lifespan: 26 years
Female sexual maturity: 10 years
Litter or clutch size: 82

Lifespan from AnAge · AnAge record · other traits from AnAge Build 15 (HAGRID 01840)

About this record

Lifespan source: AnAge build 15
Based on: an animal in the wild
Data quality: acceptable, medium sample
Licence: Free to use, subject to the Human Ageing Genomic Resources conditions of use
Status: Renamed in build 2
In the database since: build 1 (2015-07-01)
Last changed: build 2 (2026-11-13)
Earlier name: Raja rhina

Every change against the 2015 build is in the changelog download.

Composition by region

Reference (L) strand as deposited in RefSeq

Whole genome16,910 bp

Protein-coding11,411 bp

12S rRNA967 bp

16S rRNA1,675 bp

All tRNA1,549 bp

D-loop1,251 bp

Genes and regions

Select a column to sort; select a gene to compare it across species

Base composition of Caliraja rhina genes

ND1NADH dehydrogenase subunit 1	975 bp	27.7%	27.5%	12.7%	32.1%	44.82%	−0.432	+0.004
ND2NADH dehydrogenase subunit 2	1,046 bp	29.7%	27.8%	9.9%	32.5%	42.45%	−0.532	+0.033
COX1Cytochrome c oxidase subunit I	1,557 bp	25.6%	31.2%	17.0%	26.2%	43.22%	−0.212	−0.100
COX2Cytochrome c oxidase subunit II	691 bp	32.1%	28.4%	15.8%	23.7%	39.51%	−0.201	+0.062
ATP8ATP synthase F0 subunit 8	168 bp	30.4%	31.0%	8.9%	29.8%	38.69%	−0.538	−0.010
ATP6ATP synthase F0 subunit 6	684 bp	28.2%	29.7%	11.1%	31.0%	42.11%	−0.472	−0.025
COX3Cytochrome c oxidase subunit III	786 bp	26.0%	28.1%	16.4%	29.5%	45.93%	−0.285	−0.040
ND3NADH dehydrogenase subunit 3	349 bp	23.5%	31.5%	14.3%	30.7%	44.99%	−0.363	−0.146
ND4LNADH dehydrogenase subunit 4L	297 bp	24.6%	30.6%	11.8%	33.0%	44.78%	−0.474	−0.110
ND4NADH dehydrogenase subunit 4	1,381 bp	28.8%	29.1%	11.9%	30.2%	42.07%	−0.435	−0.005
ND5NADH dehydrogenase subunit 5	1,836 bp	29.8%	28.7%	12.0%	29.5%	41.45%	−0.422	+0.020
ND6NADH dehydrogenase subunit 6	519 bp	43.4%	18.3%	9.6%	28.7%	38.34%	−0.497	+0.406
CYTBCytochrome b	1,143 bp	25.9%	29.8%	13.3%	31.0%	44.27%	−0.399	−0.071
12S rRNA12S ribosomal RNA	967 bp	31.7%	25.5%	19.2%	23.5%	42.71%	−0.099	+0.108
16S rRNA16S ribosomal RNA	1,675 bp	35.0%	26.0%	17.4%	21.5%	38.93%	−0.104	+0.148
D-loopControl region (D-loop)	1,251 bp	32.6%	32.8%	12.9%	21.7%	34.61%	−0.256	−0.002

Codon usage

Counts per codon, darker is more frequent

Gene

3,809 codons in all protein-coding genes. Grouped with the Vertebrate mitochondrial code (NCBI table 2).

Phe F6.2%

UUU127

UUC109

Leu L18.0%

UUA146

UUG22

CUU127

CUC178

CUA175

CUG36

Ile I8.5%

AUU182

AUC142

Met M4.3%

AUA131

AUG32

Val V5.0%

GUU49

GUC62

GUA62

GUG18

Ser S6.7%

UCU45

UCC76

UCA78

UCG7

AGU20

AGC31

Pro P5.6%

CCU40

CCC79

CCA82

CCG11

Thr T7.7%

ACU57

ACC96

ACA135

ACG5

Ala A7.4%

GCU49

GCC142

GCA87

GCG3

Tyr Y3.2%

UAU49

UAC72

His H2.7%

CAU31

CAC70

Gln Q2.7%

CAA88

CAG13

Asn N3.8%

AAU64

AAC82

Lys K2.2%

AAA78

AAG7

Asp D1.8%

GAU32

GAC38

Glu E2.5%

GAA76

GAG21

Cys C0.6%

UGU8

UGC14

Trp W3.1%

UGA98

UGG21

Arg R1.8%

CGU15

CGC18

CGA32

CGG4

Gly G6.0%

GGU35

GGC86

GGA71

GGG36

Stop 0.2%

UAA9

UAG0

AGA0

AGG0

Genome map

38 annotated features in NC_021964.1

Hover, tap or tab to a feature to see its position, length, strand and GC content. Genes and rRNAs open their own pages.

Protein-coding13
rRNA2
tRNA22
D-loop1

Outer ring: + strand, arrows run clockwise. Inner ring: − strand.
Middle ring: GC% of each gene against the whole genome (41.4%, dashed): out is GC-richer, in is AT-richer (full height = 6 points).

All 38 features as a table

Annotated features of Caliraja rhina (NC_021964.1)
Feature	Type	Start	End	Length	Strand	GC
tRNA-Phe	Transfer RNA	1	69	69 bp	+	–
12S rRNA	Ribosomal RNA	70	1,036	967 bp	+	42.7%
tRNA-Val	Transfer RNA	1,037	1,108	72 bp	+	–
16S rRNA	Ribosomal RNA	1,109	2,783	1,675 bp	+	38.9%
tRNA-Leu (UUR)	Transfer RNA	2,784	2,858	75 bp	+	–
ND1	Protein-coding gene	2,859	3,833	975 bp	+	44.8%
tRNA-Ile	Transfer RNA	3,834	3,903	70 bp	+	–
tRNA-Gln	Transfer RNA	3,903	3,974	72 bp	−	–
tRNA-Met	Transfer RNA	3,976	4,044	69 bp	+	–
ND2	Protein-coding gene	4,045	5,090	1,046 bp	+	42.5%
tRNA-Trp	Transfer RNA	5,091	5,159	69 bp	+	–
tRNA-Ala	Transfer RNA	5,161	5,229	69 bp	−	–
tRNA-Asn	Transfer RNA	5,230	5,302	73 bp	−	–
tRNA-Cys	Transfer RNA	5,335	5,401	67 bp	−	–
tRNA-Tyr	Transfer RNA	5,402	5,471	70 bp	−	–
COX1	Protein-coding gene	5,473	7,029	1,557 bp	+	43.2%
tRNA-Ser (UCN)	Transfer RNA	7,030	7,100	71 bp	−	–
tRNA-Asp	Transfer RNA	7,101	7,169	69 bp	+	–
COX2	Protein-coding gene	7,175	7,865	691 bp	+	39.5%
tRNA-Lys	Transfer RNA	7,866	7,938	73 bp	+	–
ATP8	Protein-coding gene	7,940	8,107	168 bp	+	38.7%
ATP6	Protein-coding gene	8,098	8,781	684 bp	+	42.1%
COX3	Protein-coding gene	8,784	9,569	786 bp	+	45.9%
tRNA-Gly	Transfer RNA	9,571	9,640	70 bp	+	–
ND3	Protein-coding gene	9,641	9,989	349 bp	+	45.0%
tRNA-Arg	Transfer RNA	9,990	10,058	69 bp	+	–
ND4L	Protein-coding gene	10,059	10,355	297 bp	+	44.8%
ND4	Protein-coding gene	10,349	11,729	1,381 bp	+	42.1%
tRNA-His	Transfer RNA	11,730	11,798	69 bp	+	–
tRNA-Ser (AGY)	Transfer RNA	11,799	11,867	69 bp	+	–
tRNA-Leu (CUN)	Transfer RNA	11,868	11,939	72 bp	+	–
ND5	Protein-coding gene	11,940	13,775	1,836 bp	+	41.5%
ND6	Protein-coding gene	13,772	14,290	519 bp	−	38.3%
tRNA-Glu	Transfer RNA	14,292	14,360	69 bp	−	–
CYTB	Protein-coding gene	14,364	15,506	1,143 bp	+	44.3%
tRNA-Thr	Transfer RNA	15,510	15,582	73 bp	+	–
tRNA-Pro	Transfer RNA	15,590	15,660	71 bp	−	–
D-loop	Control region	15,660	16,910	1,251 bp	–	34.6%